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Antennaire rosée, rosy everlasting, rosy pussytoes

white margined pussytoes, white-margined everlasting, whitemargin pussytoes

Habit Gynoecious (staminate plants uncommon). Dioecious or gynoecious (staminate plants in equal frequency as pistillates or none in populations, respectively).
Plants

4–30 cm.

5–20 cm (stems sometimes stipitate-glandular, especially in dioecious diploids).

Stolons

1–7 cm.

2–7 cm (woolly).

Basal leaves

1-nerved, 8–40 × 2–10 mm, spatulate, oblanceolate, or cuneate, tips mucronate, faces usually gray-pubescent, adaxial sometimes green-glabrous.

1–3-nerved, spatulate, 15–20 × 4–6 mm, tips mucronate, abaxial faces gray-tomentose, adaxial green-glabrous (margins white woolly).

Cauline leaves

linear, 6–36 mm, usually not flagged (apices acute to subulate or with lanceolate flags).

linear, 7–16 mm, (apices acute) not flagged.

Involucres

staminate unknown; pistillate 4–10 mm.

staminate 4.5–7 mm; pistillate 5–7(–9) mm.

Corollas

staminate unknown; pistillate 2.5–6 mm.

staminate 3–5 mm; pistillate 4.5–6.5 mm.

Phyllaries

distally brown, cream, gray, green, pink, red, white, or yellow (apices acute or erose-obtuse).

(relatively wide), distally white (apices acuminate).

Heads

3–20 in corymbiform arrays.

5–8 in corymbiform arrays.

Cypselae

0.7–1.8 mm, glabrous or papillate;

pappi: staminate unknown; pistillate 3.5–6.5 mm.

0.8–2 mm, glabrous or slightly papillate;

pappi: staminate 3.5–5.5 mm; pistillate 5.5–8.5 mm.

2n

= 42, 56, (70).

= 28, 56, 84, 112, 140.

Antennaria rosea

Antennaria marginata

Phenology Flowering summer.
Habitat Moist forests, slopes and tops of ridges under Douglas fir, ponderosa pine, Engelmann spruce or Gambel oaks, openings in the forests
Elevation 1500–2900 m (4900–9500 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; TX; Mexico (Chihuahua, Coahuila)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (4 in the flora).

Antennaria rosea is the most widespread Antennaria of North America, occurring in dry to moist habitats from near sea level to the alpine zone. The A. rosea polyploid agamic complex is one of the more morphologically diverse complexes of North American Antennaria. It occurs from the western cordillera of North America from southern California, Arizona, and New Mexico north to subarctic Alaska and east to Greenland and, disjunctly, in the Canadian maritime provinces, eastern Quebec, and immediately north of and adjacent to Lake Superior (R. J. Bayer et al. 1991). Antennaria chilensis (including A. chilensis var. magellanica) is a Patagonian endemic that morphologically fits within the circumscription of A. rosea and may well be an amphitropical disjunct member of the complex.

Antennaria rosea is taxonomically confusing; it includes agamospermous microspecies that have been recognized as distinct taxonomic species. Morphometric and isozyme analyses have demonstrated that the primary source of morphologic variability in the complex derives from six sexually reproducing progenitors, A. aromatica, A. corymbosa, A. pulchella, A. microphylla, A. racemosa, and A. umbrinella (R. J. Bayer 1989b, 1990b, 1990c). Additionally, three other sexually reproducing species, A. marginata, A. suffrutescens, and A. rosulata, may have contributed to the genetic complexity of the A. rosea complex (Bayer 1990b). Here, four reasonably distinct subspecies are recognized within the complex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Antennaria marginata has rims of white hairs (from the abaxial faces) around its adaxially glabrous leaves. It has both dioecious and gynoecious populations and cytotypes ranging from diploid to decaploid (R. J. Bayer and G. L. Stebbins 1987). It is probably a primary sexual progenitor of the A. parvifolia polyploid complex; the two taxa sometimes overlap morphologically; they differ in induments of basal leaves. Antennaria marginata may also be a contributor to the parentage of the A. howellii and A. rosea agamic complexes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 20–40 mm; phyllaries distally usually green, pink, red or white, seldom brown
subsp. rosea
1. Basal leaves 8–20 mm; phyllaries distally brown, cream, gray, green, pink, red, white, or yellow
→ 2
2. Pistillate: involucres 4–6.5 mm, corollas 2.5–4, pappi 3.5–5; cauline leaves 6–20 mm (tips subulate); phyllaries usually distally brown, sometimes cream, gray, or yellow
subsp. confinis
2. Pistillate: involucres 6.5–10 mm, corollas 3.5–6 mm, pappi 5–6.5 mm; cauline leaves 6–19 or 9–26 mm (tips sometimes with flat, lanceolate scarious appendages); phyllaries distally brown, green, pink, red, or white
→ 3
3. Plants 19–30 cm; cauline leaves 9–26 mm (proximalmost usually 19+ mm); heads usu- ally 6–12
subsp. arida
3. Plants 4–17 cm; cauline leaves 6–19 mm (proximalmost usually less than 19 mm); heads usually 3–5
subsp. pulvinata
Source FNA vol. 19, p. 408. FNA vol. 19, p. 405.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. rosea subsp. arida, A. rosea subsp. confinis, A. rosea subsp. pulvinata, A. rosea subsp. rosea
Synonyms A. dioica var. marginata, A. fendleri, A. marginata var. glandulifera, A. peramoena
Name authority Greene: Pittonia 3: 281. (1898) Greene: Pittonia 3: 290. (1898)
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