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Antennaire rosée, rosy everlasting, rosy pussytoes

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Gynoecious (staminate plants uncommon). Gynoecious (staminate plants very uncommon).
Plants

4–30 cm.

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

1–7 cm.

1–9(–12) cm.

Basal leaves

1-nerved, 8–40 × 2–10 mm, spatulate, oblanceolate, or cuneate, tips mucronate, faces usually gray-pubescent, adaxial sometimes green-glabrous.

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

linear, 6–36 mm, usually not flagged (apices acute to subulate or with lanceolate flags).

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate unknown; pistillate 4–10 mm.

staminate 6–6.5 mm; pistillate 6–11 mm.

Corollas

staminate unknown; pistillate 2.5–6 mm.

staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.

Phyllaries

distally brown, cream, gray, green, pink, red, white, or yellow (apices acute or erose-obtuse).

(bases sometimes rose) distally white, cream, or light brown.

Heads

3–20 in corymbiform arrays.

3–15 in corymbiform arrays.

Cypselae

0.7–1.8 mm, glabrous or papillate;

pappi: staminate unknown; pistillate 3.5–6.5 mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.

2n

= 42, 56, (70).

= 56, 84, 140 (under A. neodioica).

Antennaria rosea

Antennaria howellii

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (4 in the flora).

Antennaria rosea is the most widespread Antennaria of North America, occurring in dry to moist habitats from near sea level to the alpine zone. The A. rosea polyploid agamic complex is one of the more morphologically diverse complexes of North American Antennaria. It occurs from the western cordillera of North America from southern California, Arizona, and New Mexico north to subarctic Alaska and east to Greenland and, disjunctly, in the Canadian maritime provinces, eastern Quebec, and immediately north of and adjacent to Lake Superior (R. J. Bayer et al. 1991). Antennaria chilensis (including A. chilensis var. magellanica) is a Patagonian endemic that morphologically fits within the circumscription of A. rosea and may well be an amphitropical disjunct member of the complex.

Antennaria rosea is taxonomically confusing; it includes agamospermous microspecies that have been recognized as distinct taxonomic species. Morphometric and isozyme analyses have demonstrated that the primary source of morphologic variability in the complex derives from six sexually reproducing progenitors, A. aromatica, A. corymbosa, A. pulchella, A. microphylla, A. racemosa, and A. umbrinella (R. J. Bayer 1989b, 1990b, 1990c). Additionally, three other sexually reproducing species, A. marginata, A. suffrutescens, and A. rosulata, may have contributed to the genetic complexity of the A. rosea complex (Bayer 1990b). Here, four reasonably distinct subspecies are recognized within the complex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 20–40 mm; phyllaries distally usually green, pink, red or white, seldom brown
subsp. rosea
1. Basal leaves 8–20 mm; phyllaries distally brown, cream, gray, green, pink, red, white, or yellow
→ 2
2. Pistillate: involucres 4–6.5 mm, corollas 2.5–4, pappi 3.5–5; cauline leaves 6–20 mm (tips subulate); phyllaries usually distally brown, sometimes cream, gray, or yellow
subsp. confinis
2. Pistillate: involucres 6.5–10 mm, corollas 3.5–6 mm, pappi 5–6.5 mm; cauline leaves 6–19 or 9–26 mm (tips sometimes with flat, lanceolate scarious appendages); phyllaries distally brown, green, pink, red, or white
→ 3
3. Plants 19–30 cm; cauline leaves 9–26 mm (proximalmost usually 19+ mm); heads usu- ally 6–12
subsp. arida
3. Plants 4–17 cm; cauline leaves 6–19 mm (proximalmost usually less than 19 mm); heads usually 3–5
subsp. pulvinata
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 408. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. rosea subsp. arida, A. rosea subsp. confinis, A. rosea subsp. pulvinata, A. rosea subsp. rosea
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Greene: Pittonia 3: 281. (1898) Greene: Pittonia 3: 174. (1897)
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