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Hooker's pussy-toes, raceme pussytoes, racemose pussytoes, slender pussy-toes

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Dioecious. Gynoecious (staminate plants very uncommon).
Plants

12–50 cm (stems stipitate-glandular distally).

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

3–8 cm.

1–9(–12) cm.

Basal leaves

3-nerved, elliptic to oblong, 30–100 × 10–40 mm, tips mucronate, abaxially tomentose, adaxially glabrous.

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

linear, 10–30 mm, not flagged (apices obtuse to acute).

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate 4–8 mm; pistillate 7–9 mm.

staminate 6–6.5 mm; pistillate 6–11 mm.

Corollas

staminate 3–4 mm; pistillate 3–4 mm.

staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.

Phyllaries

(relatively wide) distally white or light brown (apices blunt).

(bases sometimes rose) distally white, cream, or light brown.

Heads

3–12 in loose, racemiform to paniculiform arrays.

3–15 in corymbiform arrays.

Cypselae

1–1.5 mm, glabrous or slightly papillate;

pappi: staminate 3–4.5 mm; pistillate 4.5–7 mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.

2n

= 28.

= 56, 84, 140 (under A. neodioica).

Antennaria racemosa

Antennaria howellii

Phenology Flowering summer.
Habitat Moist, cool, montane and subalpine coniferous forests and roadcuts in forests
Elevation 1200–3000 m (3900–9800 ft)
Distribution
from FNA
CA; ID; MT; OR; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria racemosa is characterized by adaxially glabrous basal leaves and open, racemiform to paniculiform arrays of heads (R. J. Bayer 1985b). The young leaves have a slight odor of citronella when crushed. Antennaria racemosa has a pivotal sexual genome of the Catipes group and has contributed to the origin of clones in the A. howellii, A. parlinii, and A. rosea polyploid agamic complexes (Bayer 1985, 1985b, 1990b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 401. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. petasites, A. piperi A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Hooker: Fl. Bor.-Amer. 1: 330. (1834) Greene: Pittonia 3: 174. (1897)
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