The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

little-leaf pussytoes, Nuttall's pussytoes, small-leaf pussytoes

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Dioecious or gynoecious (staminate plants uncommon or in equal frequency as pistillates, respectively). Gynoecious (staminate plants very uncommon).
Plants

2–8(–15) cm.

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

1–6 cm.

1–9(–12) cm.

Basal leaves

1-nerved, narrowly spatulate to spatulate or oblanceolate, 8–35 × 2–15 mm, tips mucronate, faces gray-tomentose.

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

linear to narrowly oblanceolate, 8–20 mm, not flagged (apices acute).

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate 5.5–7.5 mm;

pistillate 8–10(–15) mm (gynoecious), 7–7.2 mm (dioecious).

staminate 6–6.5 mm;

pistillate 6–11 mm.

Corollas

staminate 3.5–4.5 mm;

pistillate 5–8 mm.

staminate 3–4 mm;

pistillate 3.5–6.5(–8) mm.

Phyllaries

distally white, pink, green, red, or brown.

(bases sometimes rose) distally white, cream, or light brown.

Heads

2–7 in corymbiform arrays.

3–15 in corymbiform arrays.

Cypselae

1–1.8 mm, glabrous or minutely papillate;

pappi: staminate 4–5.5 mm;

pistillate 6.5–9 mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm;

pistillate 5.5–9 mm.

2n

= 56, 84, 112, 140.

= 56, 84, 140 (under A. neodioica).

Antennaria parvifolia

Antennaria howellii

Phenology Flowering late spring–summer.
Habitat Prairies, pastures, roadsides, mountain parks, open deciduous woods, and drier coniferous forests, usually ponderosa or lodgepole pine
Elevation 100–3400 m (300–11200 ft)
Distribution
from FNA
AZ; CO; IA; ID; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; MB; ON; SK; Okla (expected in panhandle); Wis (expected); Mexico (Chihuahua, Nuevo León)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria parvifolia is a widespread, polyploid complex of sexual (dioecious) and asexual (gynoecious) populations (G. L. Stebbins 1932b; R. J. Bayer and Stebbins 1987). Although variable morphologically, no infraspecific taxa seem warranted at this time. Sexual (dioecious) populations are known primarily from New Mexico and Colorado; apomictic plants occur throughout the range of the species. Probable sexual diploid/tetraploid progenitors of the A. parvifolia complex include A. dioica, A. marginata, A. neglecta, and A. pulchella/A. media. Antennaria parvifolia is characterized by relatively short stature and relatively small numbers of relatively large heads. The epithet parvifolia has been rendered as “parviflora” in floras, e.g., key in Great Plains Flora Association (1986); E. H. Moss (1959); H. J. Scoggan (1978–1979, part 4). In some floras, A. parvifolia has been confused with A. microphylla; the two are probably not closely related.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 406. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. aprica, A. aprica var. aureola, A. aprica var. minuscula, A. aureola, A. dioica var. parvifolia, A. holmii, A. latisquamea, A. minuscula, A. recurva, A. rhodantha A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 406. (1841) Greene: Pittonia 3: 174. (1897)
Web links