Antennaria parvifolia |
Antennaria howellii |
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little-leaf pussytoes, Nuttall's pussytoes, small-leaf pussytoes |
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes |
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Habit | Dioecious or gynoecious (staminate plants uncommon or in equal frequency as pistillates, respectively). | Gynoecious (staminate plants very uncommon). | ||||||||||||
Plants | 2–8(–15) cm. |
(6–)8–35 cm (stems sometimes stipitate-glandular). |
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Stolons | 1–6 cm. |
1–9(–12) cm. |
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Basal leaves | 1-nerved, narrowly spatulate to spatulate or oblanceolate, 8–35 × 2–15 mm, tips mucronate, faces gray-tomentose. |
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent. |
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Cauline leaves | linear to narrowly oblanceolate, 8–20 mm, not flagged (apices acute). |
linear, 8–40 mm, distal sometimes flagged (apices acute). |
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Involucres | staminate 5.5–7.5 mm; pistillate 8–10(–15) mm (gynoecious), 7–7.2 mm (dioecious). |
staminate 6–6.5 mm; pistillate 6–11 mm. |
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Corollas | staminate 3.5–4.5 mm; pistillate 5–8 mm. |
staminate 3–4 mm; pistillate 3.5–6.5(–8) mm. |
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Phyllaries | distally white, pink, green, red, or brown. |
(bases sometimes rose) distally white, cream, or light brown. |
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Heads | 2–7 in corymbiform arrays. |
3–15 in corymbiform arrays. |
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Cypselae | 1–1.8 mm, glabrous or minutely papillate; pappi: staminate 4–5.5 mm; pistillate 6.5–9 mm. |
0.8–2 mm, ± papillate; pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm. |
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2n | = 56, 84, 112, 140. |
= 56, 84, 140 (under A. neodioica). |
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Antennaria parvifolia |
Antennaria howellii |
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Phenology | Flowering late spring–summer. | |||||||||||||
Habitat | Prairies, pastures, roadsides, mountain parks, open deciduous woods, and drier coniferous forests, usually ponderosa or lodgepole pine | |||||||||||||
Elevation | 100–3400 m (300–11200 ft) | |||||||||||||
Distribution |
AZ; CO; IA; ID; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; MB; ON; SK; Okla (expected in panhandle); Wis (expected); Mexico (Chihuahua, Nuevo León)
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CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
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Discussion | Antennaria parvifolia is a widespread, polyploid complex of sexual (dioecious) and asexual (gynoecious) populations (G. L. Stebbins 1932b; R. J. Bayer and Stebbins 1987). Although variable morphologically, no infraspecific taxa seem warranted at this time. Sexual (dioecious) populations are known primarily from New Mexico and Colorado; apomictic plants occur throughout the range of the species. Probable sexual diploid/tetraploid progenitors of the A. parvifolia complex include A. dioica, A. marginata, A. neglecta, and A. pulchella/A. media. Antennaria parvifolia is characterized by relatively short stature and relatively small numbers of relatively large heads. The epithet parvifolia has been rendered as “parviflora” in floras, e.g., key in Great Plains Flora Association (1986); E. H. Moss (1959); H. J. Scoggan (1978–1979, part 4). In some floras, A. parvifolia has been confused with A. microphylla; the two are probably not closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (4 in the flora). The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 406. | FNA vol. 19, p. 403. | ||||||||||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | A. aprica, A. aprica var. aureola, A. aprica var. minuscula, A. aureola, A. dioica var. parvifolia, A. holmii, A. latisquamea, A. minuscula, A. recurva, A. rhodantha | A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii | ||||||||||||
Name authority | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 406. (1841) | Greene: Pittonia 3: 174. (1897) | ||||||||||||
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