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Antennaire négligée, field pussytoes

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Dioecious. Gynoecious (staminate plants very uncommon).
Plants

4–25 cm.

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

2.5–18 cm.

1–9(–12) cm.

Basal leaves

1-nerved, narrowly spatulate to cuneate-oblanceolate, 15–65 × 6–18 mm, tips mucronate, faces abaxially tomentose, adaxially gray-pubescent (green-glabrescent with age).

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

linear, 1.5–25 mm, distal flagged.

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate 4–7 mm; pistillate 6–10 mm.

staminate 6–6.5 mm; pistillate 6–11 mm.

Corollas

staminate 2.7–5 mm; pistillate 4.5–6.5(–7) mm.

staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.

Phyllaries

distally white.

(bases sometimes rose) distally white, cream, or light brown.

Heads

(1–)2–8 in corymbiform to spiciform or racemiform arrays.

3–15 in corymbiform arrays.

Cypselae

0.9–1.4 mm, minutely papillate;

pappi: staminate 3.5–6.5 mm; pistillate 6–8.5(–9.5) mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.

2n

= 28.

= 56, 84, 140 (under A. neodioica).

Antennaria neglecta

Antennaria howellii

Phenology Flowering early–mid spring.
Habitat Plains, grasslands, pastures, and open woodlands
Elevation 0–2500 m (0–8200 ft)
Distribution
from FNA
AR; CO; CT; DE; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SD; VA; VT; WI; WV; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria neglecta is a sexual progenitor of both the A. howellii and A. parvifolia polyploid complexes and has one of the more widespread ranges among the amphimictic species in the genus in North America. Amphimicts generally have small ranges compared to those of the polyploid agamic complexes derived from them. Characteristic features of A. neglecta are its lashlike stolons that bear reduced leaves (except at the ends), flags on the distal cauline leaves, and basal leaves that are green-glabrescent with age (R. J. Bayer and G. L. Stebbins 1982).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 403. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. athabascensis, A. campestris, A. campestris var. athabascensis, A. chelonica, A. erosa, A. howellii var. athabascensis, A. howellii var. campestris, A. longifolia, A. lunellii, A. nebrascensis, A. neglecta var. athabascensis, A. neglecta var. campestris, A. neglecta var. simplex, A. parvula, A. wilsonii A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Greene: Pittonia 3: 173. (1897) Greene: Pittonia 3: 174. (1897)
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