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little-leaf pussytoes, pink pussytoes, rosy pussytoes, small pussytoes, white pussytoes

Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes

Habit Dioecious. Gynoecious (staminate plants very uncommon).
Plants

9–30 cm (stems stipitate-glandular distally).

(6–)8–35 cm (stems sometimes stipitate-glandular).

Stolons

1–5 cm.

1–9(–12) cm.

Basal leaves

1-nerved, spatulate, 6–16 × 2–6 mm, tips mucronate, faces silvery gray-pubescent.

1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.

Cauline leaves

linear, 5–25 mm, not flagged (apices acute).

linear, 8–40 mm, distal sometimes flagged (apices acute).

Involucres

staminate 5–6.5 mm; pistillate 5.5–7 mm.

staminate 6–6.5 mm; pistillate 6–11 mm.

Corollas

staminate 2.5–3 mm; pistillate 3–4.3 mm.

staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.

Phyllaries

distally bright white to light yellow.

(bases sometimes rose) distally white, cream, or light brown.

Heads

6–13 in corymbiform arrays.

3–15 in corymbiform arrays.

Cypselae

0.7–1.2 mm, glabrous or sparingly papillate;

pappi: staminate 3–4 mm; pistillate 3–5 mm.

0.8–2 mm, ± papillate;

pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.

2n

= 28.

= 56, 84, 140 (under A. neodioica).

Antennaria microphylla

Antennaria howellii

Phenology Flowering early–mid summer.
Habitat Moist open areas, flood plains of streams, margins of alkaline depressions, lower montane to subalpine (subarctic)
Elevation 0–3200 m (0–10500 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Antennaria microphylla is a primary sexual progenitor of the A. rosea polyploid agamic complex (R. J. Bayer 1990b). A. Cronquist (1955) included A. rosea within his circumscription of A. microphylla. It is preferable to recognize sexual diploids as distinct from their morphologically discrete hybrid apomictic derivatives. Antennaria microphylla is always dioecious and has stems distally stipitate-glandular and white phyllaries; A. rosea is always gynoecious and has stems without glandular hairs and phyllaries only occasionally white.

Some authors (A. E. Porsild 1950; E. H. Moss 1959; Porsild and W. J. Cody 1980) have recognized A. nitida as distinct; comparisons of the nomenclatural types of the two show that they are conspecific. Antennaria microphylla has allelopathic properties (G. D. Manners and D. S. Galitz 1985).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves green-glabrous adaxially
→ 2
1. Basal leaves pubescent adaxially (sometimes glabrescent with age)
→ 3
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream
subsp. canadensis
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white
subsp. howellii
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends)
subsp. neodioica
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends)
subsp. petaloidea
Source FNA vol. 19, p. 407. FNA vol. 19, p. 403.
Parent taxa Asteraceae > tribe Gnaphalieae > Antennaria Asteraceae > tribe Gnaphalieae > Antennaria
Sibling taxa
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. howellii, A. lanata, A. luzuloides, A. marginata, A. media, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
A. alpina, A. anaphaloides, A. arcuata, A. argentea, A. aromatica, A. corymbosa, A. densifolia, A. dimorpha, A. dioica, A. flagellaris, A. friesiana, A. geyeri, A. lanata, A. luzuloides, A. marginata, A. media, A. microphylla, A. monocephala, A. neglecta, A. parlinii, A. parvifolia, A. plantaginifolia, A. pulchella, A. pulcherrima, A. racemosa, A. rosea, A. rosulata, A. soliceps, A. solitaria, A. stenophylla, A. suffrutescens, A. umbrinella, A. virginica
Subordinate taxa
A. howellii subsp. canadensis, A. howellii subsp. howellii, A. howellii subsp. neodioica, A. howellii subsp. petaloidea
Synonyms A. bracteosa, A. microphylla var. solstitialis, A. nitida, A. rosea var. nitida, A. solstitialis A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii
Name authority Rydberg: Bull. Torrey Bot. Club 24: 303. (1897) Greene: Pittonia 3: 174. (1897)
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