Antennaria lanata |
Antennaria howellii |
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woolly everlasting, woolly pussy-toes |
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes |
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Habit | Dioecious. | Gynoecious (staminate plants very uncommon). | ||||||||||||
Plants | 3–20 cm (caudices branching or rhizomes stout). |
(6–)8–35 cm (stems sometimes stipitate-glandular). |
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Stolons | none. |
1–9(–12) cm. |
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Basal leaves | 3-nerved, narrowly oblanceolate, 10–60(–100) × 3–12 mm, tips acute, faces gray-woolly or tomentose. |
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent. |
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Cauline leaves | linear, 5–40 mm, mid and distal flagged. |
linear, 8–40 mm, distal sometimes flagged (apices acute). |
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Involucres | staminate 4.5–6 mm; pistillate 5–8 mm. |
staminate 6–6.5 mm; pistillate 6–11 mm. |
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Corollas | staminate 3–4.5 mm; pistillate 2.5–4 mm. |
staminate 3–4 mm; pistillate 3.5–6.5(–8) mm. |
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Phyllaries | (proximally light brown, dark brown, or olivaceous) distally whitish or light brown. |
(bases sometimes rose) distally white, cream, or light brown. |
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Heads | 3–9 in corymbiform arrays. |
3–15 in corymbiform arrays. |
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Cypselae | 1–1.6 mm, glabrous; pappi: staminate 4–5 mm; pistillate 3.5–5 mm. |
0.8–2 mm, ± papillate; pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm. |
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2n | = 28 (under A. neodioica). |
= 56, 84, 140 (under A. neodioica). |
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Antennaria lanata |
Antennaria howellii |
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Phenology | Flowering summer. | |||||||||||||
Habitat | Protected alpine and subalpine sites, gravelly or sandy soils near conifers at timberline | |||||||||||||
Elevation | 1400–3400 m (4600–11200 ft) | |||||||||||||
Distribution |
CA; ID; MT; OR; UT; WA; WY; AB; BC
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CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
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Discussion | Subspecies 4 (4 in the flora). The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 399. | FNA vol. 19, p. 403. | ||||||||||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | A. carpatica var. lanata | A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii | ||||||||||||
Name authority | (Hooker) Greene: Pittonia 3: 288. (1898) | Greene: Pittonia 3: 174. (1897) | ||||||||||||
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