Antennaria lanata |
Antennaria friesiana |
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woolly everlasting, woolly pussy-toes |
Antennaire de Fries, Fries' pussytoes |
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Habit | Dioecious. | Dioecious or gynoecious (staminate plants uncommon or in equal frequencies to pistillates, respectively). | ||||||||
Plants | 3–20 cm (caudices branching or rhizomes stout). |
7–15 cm (stems stipitate-glandular, hairs purple). |
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Stolons | none. |
0.1–4 cm. |
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Basal leaves | 3-nerved, narrowly oblanceolate, 10–60(–100) × 3–12 mm, tips acute, faces gray-woolly or tomentose. |
1-nerved, narrowly spatulate to oblanceolate, 11–30 × 2–4 mm, tips mucronate, abaxial faces tomentose, adaxial green-glabrescent to gray-pubescent. |
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Cauline leaves | linear, 5–40 mm, mid and distal flagged. |
linear, 4–20 mm, flagged. |
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Involucres | staminate 4.5–6 mm; pistillate 5–8 mm. |
staminate 4–6.5 mm; pistillate 5.5–8 mm. |
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Corollas | staminate 3–4.5 mm; pistillate 2.5–4 mm. |
staminate 2.5–3 mm; pistillate 3–4.5 mm. |
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Phyllaries | (proximally light brown, dark brown, or olivaceous) distally whitish or light brown. |
distally usually black, light brown, dark brown, or olivaceous, sometimes. |
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Heads | 3–9 in corymbiform arrays. |
2–6 in corymbiform arrays. |
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Cypselae | 1–1.6 mm, glabrous; pappi: staminate 4–5 mm; pistillate 3.5–5 mm. |
1.2–2 mm, glabrous or slightly papillate; pappi: staminate 3–4 mm; pistillate 3.5–5 mm. |
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2n | = 28 (under A. neodioica). |
= 28, 56, 63, 100+. |
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Antennaria lanata |
Antennaria friesiana |
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Phenology | Flowering summer. | |||||||||
Habitat | Protected alpine and subalpine sites, gravelly or sandy soils near conifers at timberline | |||||||||
Elevation | 1400–3400 m (4600–11200 ft) | |||||||||
Distribution |
CA; ID; MT; OR; UT; WA; WY; AB; BC
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AK; NL; NT; NU; QC; YT; Arctic North America; arctic Siberia |
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Discussion | Subspecies 3 (3 in the flora). The Antennaria friesiana complex consists of subsp. alaskana, subsp. neoalaskana, and subsp. friesiana, the former two are dioecious (sexual) phases of the latter gynoecious (asexual) form. The sexual populations are known from Alaska and cordilleran areas of northern Yukon and adjacent Northwest Territories (R. J. Bayer 1991). The apomictic phase is almost circumpolar, occurring from the central and eastern Siberian plateau eastward across the North American arctic to Greenland (Bayer). E. Hultén (1968) circumscribed a fourth subspecies, A. friesiana subsp. compacta. After studying its morphology, in the field and herbarium, it is apparent that Hultén’s taxon contains at least three incongruous entities that are probably not at all related to the other two subspecies of A. friesiana. Hultén’s subsp. compacta included A. densifolia, which is recognized as a distinct species, and A. crymophila and A. neoalaskana as taxonomic synonyms. Antennaria compacta in the strict sense and A. crymophila are perhaps hybrid apomicts and are treated here in A. alpina (see Bayer for details). Antennaria neoalaskana is treated here as a subspecies of A. friesiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 399. | FNA vol. 19, p. 412. | ||||||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | A. carpatica var. lanata | A. alpina var. friesiana | ||||||||
Name authority | (Hooker) Greene: Pittonia 3: 288. (1898) | (Trautvetter) E. Ekman: Svensk Bot. Tidskr. 22: 416. (1928) | ||||||||
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