Antennaria howellii
Antennaria marginata
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes
white margined pussytoes, white-margined everlasting, whitemargin pussytoes
(6–)8–35 cm (stems sometimes stipitate-glandular).
5–20 cm (stems sometimes stipitate-glandular, especially in dioecious diploids).
1–9(–12) cm.
2–7 cm (woolly).
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.
1–3-nerved, spatulate, 15–20 × 4–6 mm, tips mucronate, abaxial faces gray-tomentose, adaxial green-glabrous (margins white woolly).
linear, 8–40 mm, distal sometimes flagged (apices acute).
linear, 7–16 mm, (apices acute) not flagged.
staminate 6–6.5 mm; pistillate 6–11 mm.
staminate 4.5–7 mm; pistillate 5–7(–9) mm.
staminate 3–4 mm; pistillate 3.5–6.5(–8) mm.
staminate 3–5 mm; pistillate 4.5–6.5 mm.
(bases sometimes rose) distally white, cream, or light brown.
(relatively wide), distally white (apices acuminate).
3–15 in corymbiform arrays.
5–8 in corymbiform arrays.
0.8–2 mm, ± papillate;
pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm.
0.8–2 mm, glabrous or slightly papillate;
pappi: staminate 3.5–5.5 mm; pistillate 5.5–8.5 mm.
= 56, 84, 140 (under A. neodioica).
= 28, 56, 84, 112, 140.
Antennaria howellii
Antennaria marginata
Subspecies 4 (4 in the flora).
The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Antennaria marginata has rims of white hairs (from the abaxial faces) around its adaxially glabrous leaves. It has both dioecious and gynoecious populations and cytotypes ranging from diploid to decaploid (R. J. Bayer and G. L. Stebbins 1987). It is probably a primary sexual progenitor of the A. parvifolia polyploid complex; the two taxa sometimes overlap morphologically; they differ in induments of basal leaves. Antennaria marginata may also be a contributor to the parentage of the A. howellii and A. rosea agamic complexes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal leaves green-glabrous adaxially | → 2 |
1. Basal leaves pubescent adaxially (sometimes glabrescent with age) | → 3 |
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream | subsp. canadensis |
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white | subsp. howellii |
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends) | subsp. neodioica |
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends) | subsp. petaloidea |
- Local floras:
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OR, 
WA - Local Web sites:
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Flora NW, 
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MD Biodiversity, 
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PNW Herbaria, Turner Photog. 
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