Antennaria howellii
Antennaria howellii subsp. petaloidea
Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes
Antennaire pétaloïde, Howell's pussytoes, petaloid Howell's pussytoes, petaloid pussytoes, small pussytoes
(6–)8–35 cm (stems sometimes stipitate-glandular).
1–9(–12) cm.
4–9 cm (leaves along stolons smaller than in rosettes at ends).
1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent.
1(–3)-nerved, cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles), 15–45(–65) × 5–20 mm, tips mucronate, faces abaxially tomentose, adaxially gray-pubescent to floccose-glabrescent.
linear, 8–40 mm, distal sometimes flagged (apices acute).
linear, 10–35 mm, mid and distal flagged.
staminate 6–6.5 mm;
pistillate 6–11 mm.
staminate (uncommon) 6–6.5 mm;
pistillate 7–11 mm.
staminate 3–4 mm;
pistillate 3.5–6.5(–8) mm.
staminate 3–4 mm;
pistillate 4–6.5(–8) mm.
(bases sometimes rose) distally white, cream, or light brown.
distally white or cream.
3–15 in corymbiform arrays.
4–10(–15) in corymbiform arrays.
0.8–2 mm, ± papillate;
pappi: staminate 4–4.5 mm;
pistillate 5.5–9 mm.
0.8–1.7 mm, minutely papillate;
pappi: staminate 4–4.5 mm;
pistillate 5.5–8 mm.
= 56, 84, 140 (under A. neodioica).
= 56, 84 (under A. neodioica).
Antennaria howellii
Antennaria howellii subsp. petaloidea
Subspecies 4 (4 in the flora).
The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies petaloidea is most common in the eastern half of the range of Antennaria howellii and is frequent as far west as British Columbia and Washington. Its primary sexual progenitors include A. plantaginifolia and A. neglecta (R. J. Bayer 1985).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal leaves green-glabrous adaxially | → 2 |
1. Basal leaves pubescent adaxially (sometimes glabrescent with age) | → 3 |
2. Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream | subsp. canadensis |
2. Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white | subsp. howellii |
3. Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends) | subsp. neodioica |
3. Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends) | subsp. petaloidea |
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