Antennaria howellii |
Antennaria aromatica |
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Antennaire de Howell, everlasting pussytoes, Howell's pussytoes, small pussytoes |
aromatic pussytoes, scented pussytoes |
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Habit | Gynoecious (staminate plants very uncommon). | Dioecious. | ||||||||||||
Plants | (6–)8–35 cm (stems sometimes stipitate-glandular). |
2–7 cm (stems stipitate-glandular). |
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Stolons | 1–9(–12) cm. |
0.5–2.5 cm. |
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Basal leaves | 1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48(–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent. |
1-nerved, usually cuneate-spatulate, sometimes oblanceolate, 5–16 × 3–10 mm, tips mucronate, faces gray-pubescent (and stipitate-glandular; fresh leaves citronella scented). |
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Cauline leaves | linear, 8–40 mm, distal sometimes flagged (apices acute). |
linear, 3–14 mm, not flagged (apices acute). |
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Involucres | staminate 6–6.5 mm; pistillate 6–11 mm. |
staminate 4.5–6.5 mm; pistillate 5–7(–9) mm. |
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Corollas | staminate 3–4 mm; pistillate 3.5–6.5(–8) mm. |
staminate 2.5–3 mm; pistillate 3.5–4.5 mm. |
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Phyllaries | (bases sometimes rose) distally white, cream, or light brown. |
distally light brown, dark brown, or olivaceous. |
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Heads | 3–15 in corymbiform arrays. |
borne singly or 2–5 in corymbiform arrays. |
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Cypselae | 0.8–2 mm, ± papillate; pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm. |
0.9–2 mm, sparingly papillate; pappi: staminate 3–4 mm; pistillate 4.5–5.5 mm. |
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2n | = 56, 84, 140 (under A. neodioica). |
= 28, 56, 84. |
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Antennaria howellii |
Antennaria aromatica |
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Phenology | Flowering mid summer. | |||||||||||||
Habitat | Subalpine limestone talus | |||||||||||||
Elevation | 1600–3000 m (5200–9800 ft) | |||||||||||||
Distribution |
CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT
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ID; MT; WY; AB
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Discussion | Subspecies 4 (4 in the flora). The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Known only from the northern Rockies, Antennaria aromatica is characterized by glandulosity, cuneate leaves, and odor of citronella in crushed leaves of living material. It is most closely related to A. densifolia of the Northwest Territories and Yukon (R. J. Bayer 1989c). Some collections of pistillate plants from Colorado and other areas of the Rockies superficially resemble A. aromatica and undoubtedly have A. aromatica in their parentage. They are non-glandular and odorless and are closer to the type of A. pulvinata, which is included in the circumscription of A. rosea, as A. rosea subsp. pulvinata (Bayer). Antennaria aromatica is a sexual progenitor of the A. rosea and A. alpina polyploid complexes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 403. | FNA vol. 19, p. 408. | ||||||||||||
Parent taxa | Asteraceae > tribe Gnaphalieae > Antennaria | Asteraceae > tribe Gnaphalieae > Antennaria | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | A. neglecta subsp. howellii, A. neglecta var. howellii, A. neodioica subsp. howellii | |||||||||||||
Name authority | Greene: Pittonia 3: 174. (1897) | Evert: Madroño 31: 109, fig. 1. (1984) | ||||||||||||
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