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heartleaf madeiravine, Madeira-vine, mignonette vine

Madeira-vine, madieravine

Roots

fibrous or tuberous-thickened, fleshy.

Stems

twining to 5 m, often producing single or clustered axillary tubers.

herbaceous, not ridged, to 3 cm diam.;

epidermis scaly and exfoliating with age;

aerial tubers axillary, basal, or absent.

Leaves

petiole 6–12 mm;

blade ovate to orbicular, 2–10 × 1–7 cm, base of larger leaf blades proximal to inflorescences cordate, base of small distal leaf blades tapering, apex obtuse to acute.

petiolate to subsessile;

blade orbicular to elliptic, ovate, or cordate, base cuneate to truncate or cordate, margins entire, apex acute to acuminate or obtuse.

Inflorescences

racemes or in branched panicles of racemes, 10–35(–60) cm;

single bract subtending each pedicel triangular-lanceolate, 1–2 × 0.3–0.4 mm; paired bracts subtending each flower persistent, triangular to obtuse, 0.6–1 × 0.5–0.6 mm, basally connate into cup.

indeterminate, axillary or terminal, racemose or clusters of racemes;

bracts persistent or deciduous, connate or distinct, membranous to succulent.

Flowers

bisexual, usually functionally staminate;

sepals basally adnate to petals, cream-white, not winged in fruit, ovate to elliptic, 1.2–2.3 × 1.1–2 mm, apex obtuse;

petals basally connate, cream-white, ovate to elliptic, 2.1–3 × 1.4–2 mm, apex obtuse, spreading at anthesis;

stamens fleshy;

filaments basally connate and dilated, 1.9–3.5 mm;

anthers early deciduous, 0.7–0.9 mm;

pistils 0.4–0.6 mm;

styles 1–1.5 mm, basally connate for 1/2–2/3 their length;

stigmas clavate to capitate;

pedicel 1–3 mm.

bisexual or functionally unisexual, fragrant, pedicellate;

sepals distinct or basally connate, adaxially rounded to keeled, elliptic to ovate or obovate, basally adnate to petals, apex acute or obtuse;

petals 5, distinct ± to base, erect to spreading or reflexed at anthesis;

stamens opposite petals;

filaments recurved in bud, flattened and proximally dilated;

anthers versatile, dehiscence longitudinal;

pollen pantoporate, spheric;

pistils globose to ovoid;

stigmas clavate to capitate or 2-lobed.

Seeds

erect, laterally flattened to nearly globose.

Utricles

rarely producing viable seeds, style bases persistent, globose, 0.8–1.1 mm.

partly to completely enclosed in persistent, dry perianth, wing or not.

x

= 12.

2n

= 24.

Anredera cordifolia

Anredera

Phenology Flowering late summer–fall.
Habitat Disturbed areas, fencerows, roadsides
Elevation 0-500 m (0-1600 ft)
Distribution
from FNA
CA; FL; LA; TX; Mexico; Central America; West Indies; Eurasia; Africa; Pacific Islands; Australia; native to South America [Introduced in North America]
[BONAP county map]
from USDA
North America; Central America; South America; West Indies [Introduced in Eurasia, Africa, Pacific Islands, Australia]
[BONAP county map]
Discussion

Anredera cordifolia is sometimes cultivated as an ornamental, and it escapes and naturalizes in subtropical to subtemperate regions (C. R. Sperling 1987). In many floristic treatments, it has been confused with A. baselloides (Kunth) Baillon, a species restricted to Ecuador and Peru. Fruit- and seed-set are rare in cultivated material; tubers are the main means by which Madeira- vine is propagated (C. R. Sperling 1987).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 12 (2 in the flora).

Anredera cordifolia (Tenore) Steenis has been shown to have anti-inflammatory, anti-ulcer, and liver-protective effects in rats (as Boussingaultia gracilis; Lin C. C. et al. 1994; Lin W. C. et al. 1995, 1996). Anredera vesicaria (Lamarck) Gaertner has been used in Mexico to treat broken bones and flesh wounds (as A. scandens; R. Mata 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Larger leaves proximal to inflorescence cordate at base; pedicel bracts basally connate, per- sisting at apex of pedicel; sepals not winged in fruit; axillary tubers often produced
A. cordifolia
1. Larger leaves proximal to inflorescence tapering at base; pedicel bracts distinct, deciduous; sepals winged in fruit; without axillary tubers
A. vesicaria
Source FNA vol. 4, p. 507. FNA vol. 4, p. 506.
Parent taxa Basellaceae > Anredera Basellaceae
Sibling taxa
A. vesicaria
Subordinate taxa
A. cordifolia, A. vesicaria
Synonyms Boussingaultia cordifolia, Boussingaultia gracilis Boussingaultia
Name authority (Tenore) Steenis: Fl. Males. Ser. 1, 5: 303. (1957) Jussieu: Gen. Pl., 84. (1789)
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