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Habit Plants small, filiform to wiry, dark brownish green. Plants small, in loose mats, dark when dry, bright green when moist.

0.5–1.5 cm, 0.3–0.5 mm thick when dry, rarely branched, primary branches prostrate;

central strand cells not differentiated;

pseudoparaphyllia absent;

rhizoids few.

1.5–3 cm, 0.5–1 mm thick when dry, somewhat densely branched, primary branches erect to ascending, julaceous when dry;

central strand cells differentiated;

pseudoparaphyllia absent;

rhizoids few.

Branch leaves

julaceous when dry, spreading, not complanate when moist, delicate, narrowly ligulate to tapering, 0.5–0.9 mm;

base narrowly decurrent;

margins plane, crenulate toward apex by prominent papillae;

apex narrowly obtuse to acuminate, often broken off;

costa weak, ending before mid leaf, obscured by laminal cells almost throughout, pellucid proximally, abaxial costa cells smooth;

basal laminal cells few, pellucid, smooth, region not reaching margin;

medial cells round, 4 µm, papillae many, unbranched.

slightly crisped, incurved when dry, secund, reflexed, not complanate when moist, ligulate, 1.2–1.8(–2.1) mm;

base ovate;

margins plane, not papillose-crenulate at insertion;

apex obtuse to rounded, often broken off;

costa moderately strong, ending much before apex, obscured by laminal cells distally, light green, abaxial costa cells papillose, papillae large, in rows;

basal laminal cells many, pellucid, smooth, region reaching margin, to 1/2 basal portion of lamina or beyond;

medial and distal cells hexagonal, 6–11 µm, papillae many, branched.


rare, on terminal branches, leaves oblong, apex acuminate, laminal cells with 1 or 2 papillae per lumen.

on terminal branches, beyond distalmost branching points, near apices, leaves with laminal cells papillose.




Anomodon tristis

Anomodon thraustus

Habitat Bark of trees, deciduous forests Bark of trees, deciduous forests
Elevation moderate to high elevations moderate elevations
from FNA
AL; AR; AZ; CT; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; VT; WI; WV; HI; NB; NS; ON; QC; Mexico (Jalisco, Nuevo León, Sonora, Veracruz); Central America (Costa Rica); South America (Bolivia); Europe; Asia
from FNA
NC; NJ; NY; Mexico (Hidalgo, Jalisco, San Luis Potosí); Asia

Although the degree to which the apex breaks off is variable within Anomodon tristis, the character allows for easy identification of this species and others in sect. Haplohymenium. However, other taxa outside the section also present this feature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Anomodon thraustus resembles A. minor but can be distinguished by the more delicate habit, thinner branches, differentiated central strand of smaller cells, and incurved or crisped leaves that become widely spreading when moist rather than complanate. The stems are sometimes irregularly pinnate. Leaves of A. thraustus are narrow with broken apices, similar to those of A. tristis, and the costa is obscured by laminal cells distally. The delicate, narrowly ligulate distal leaf of A. thraustus is distinct; that of A. minor is wider and shorter. Anomodon thraustus has been considered an East Asian species, but isolated records from central Mexico exist (L. J. Gier 1980). All North American (and most Mexican) specimens of this taxon have been misidentified as A. minor (and could also be mistaken for A. tristis). Although it is a rare species, the presence of A. thraustus in the New World has been overlooked. The disjunct distribution of A. thraustus parallels that of A. minor, although in the New World A. thraustus is somewhat more southerly in range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 633. FNA vol. 28, p. 633.
Parent taxa Anomodontaceae > Anomodon Anomodontaceae > Anomodon
Sibling taxa
A. attenuatus, A. longifolius, A. minor, A. rostratus, A. rugelii, A. thraustus, A. viticulosus
A. attenuatus, A. longifolius, A. minor, A. rostratus, A. rugelii, A. tristis, A. viticulosus
Synonyms Leskea tristis, Haplohymenium triste, Hypnum triste
Name authority (Cesati) Sullivant & Lesquereux: in W. S. Sullivant, Musc. Hepat. U.S., 241. (1856) Müller Hal.: Nuovo Giorn. Bot. Ital., n. s. 5: 207. (1898)
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