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anomodon moss, poodle moss

Habit Plants large, in dense mats, yellowish green. Plants small to large, in dense or loose mats, glaucous, green, brown, or yellowish brown, dull.
Stem(s)

1.5–3 cm, 0.8–1 mm thick when dry, profusely branched, irregularly pinnate, primary branches prostrate to arcuate, mostly attenuate, secondary branches attenuate at apices;

central strand cells not differentiated;

pseudoparaphyllia absent;

rhizoids somewhat abundant.

and branch leaves differentiated.;

stem leaves scalelike, minute;

apex acute-acuminate to rounded;

costa single, long, ending below apex, thick, usually pellucid, or double and short;

laminal cells short.

Branch leaves

appressed when dry, complanate when moist, ligulate, slightly narrowed mid leaf, 1.2–2(–2.2) mm;

base broadly decurrent;

margins plane, entire from base to apex (cells moderately mammillose at insertion), sometimes denticulate near apex;

apex acute, sometimes obtuse or slightly apiculate, intact;

costa strong, sharply ending a few cells before apex, sometimes fading in youngest leaves, not obscured by laminal cells distally, pellucid, abaxial costa cells smooth;

basal laminal cells hyaline or sometimes chlorophyllose, papillae few, region sometimes extending more than 1/2 length of leaf base;

distal cells quadrate, 6–8 µm, papillae many, branched.

with costa single, ending sharply at or near apex, pellucid.

Seta

1–1.5 cm.

dark to light reddish or brown, flexuose.

Sexual condition

dioicous.

Capsule

oblong, urn 1.6–2.8 mm;

stomata at base;

annulus not differentiated;

operculum obliquely short-rostrate, 1.3 mm;

exostome teeth irregular, 0.3 mm, striolate proximally, trabeculate and ± finely papillose towards apex;

endostome basal membrane 3 or 4 cells high, segments well developed.

erect, exserted;

operculum conic to obliquely short-rostrate;

exostome whitish yellow to pale brown, often striolate at base and papillose.

Spores

10–13 µm, densely papillose.

Perichaetia

never beyond last branching points, leaves abruptly narrowed, costa to beyond mid leaf, ending near apex in most interior leaves, laminal cells smooth.

Anomodon attenuatus

Anomodontaceae

Phenology Capsules mature early-mid fall.
Habitat Tree bark, at base, soil, rock
Elevation low to moderate elevations
Distribution
from FNA
AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; MB; NB; NF; NS; ON; PE; QC; Mexico; West Indies (Cuba, Hispaniola, Jamaica); Central America (Belize, Guatemala); Europe; Asia (India, e Russia, Turkey)
[WildflowerSearch map]
North America; Mexico; Central America; South America; West Indies; Europe; e Asia; Africa; Pacific Islands; Australia; circumboreal areas
Discussion

Anomodon attenuatus is a rather polymorphic species producing abundant sporophytes in North America. The attenuate branches, sometimes arcuate and seldom ascending, are distinctive. However, this character is sometimes lacking; in some specimens the branch apices become slightly capitate and incurved. Robust plants of A. attenuatus might be mistaken for those of A. viticulosus as both have acute leaf apices. They can be distinguished by their branch morphology: prostrate and complanate in A. attenuatus, ascending and robust in A. viticulosus. Also, the leaf shape is different in both species: lanceolate and with no constriction in A. attenuatus, broadly ligulate, abruptly narrowed from a broadly ovate base in A. viticulosus. The costa in A. attenuatus also lacks the thick, aligned papillae on the dorsal surface. A consistent character in fertile plants is the absence of gametangia beyond the most recent branching points (they are never formed in the youngest branches). The branching pattern is often of several orders of successive branching, in stepwise fashion; the endostome is papillose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 4, species ca. 20 (2 genera, 9 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branch leaves broadly ovate to lanceolate, ± abruptly narrowed mid leaf; laminal cells with papillae 1 or many.
Anomodon
1. Branch leaves ovate-lanceolate, gradually tapered from base to apex; laminal cells smooth.
Herpetineuron
Source FNA vol. 28, p. 634. FNA vol. 28, p. 629. Author: Íñigo Granzow-de la Cerda.
Parent taxa Anomodontaceae > Anomodon
Sibling taxa
A. longifolius, A. minor, A. rostratus, A. rugelii, A. thraustus, A. tristis, A. viticulosus
Subordinate taxa
Anomodon, Herpetineuron
Synonyms Leskea attenuata, Hypnum attenuatum
Name authority (Hedwig) Huebener: Muscol. Germ., 562. (1833) Kindberg
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