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anomodon moss, poodle moss

Habit Plants large, in dense mats, yellowish green. Plants small, filiform to wiry, dark brownish green.
Stems

1.5–3 cm, 0.8–1 mm thick when dry, profusely branched, irregularly pinnate, primary branches prostrate to arcuate, mostly attenuate, secondary branches attenuate at apices;

central strand cells not differentiated;

pseudoparaphyllia absent;

rhizoids somewhat abundant.

0.5–1.5 cm, 0.3–0.5 mm thick when dry, rarely branched, primary branches prostrate;

central strand cells not differentiated;

pseudoparaphyllia absent;

rhizoids few.

Branch leaves

appressed when dry, complanate when moist, ligulate, slightly narrowed mid leaf, 1.2–2(–2.2) mm;

base broadly decurrent;

margins plane, entire from base to apex (cells moderately mammillose at insertion), sometimes denticulate near apex;

apex acute, sometimes obtuse or slightly apiculate, intact;

costa strong, sharply ending a few cells before apex, sometimes fading in youngest leaves, not obscured by laminal cells distally, pellucid, abaxial costa cells smooth;

basal laminal cells hyaline or sometimes chlorophyllose, papillae few, region sometimes extending more than 1/2 length of leaf base;

distal cells quadrate, 6–8 µm, papillae many, branched.

julaceous when dry, spreading, not complanate when moist, delicate, narrowly ligulate to tapering, 0.5–0.9 mm;

base narrowly decurrent;

margins plane, crenulate toward apex by prominent papillae;

apex narrowly obtuse to acuminate, often broken off;

costa weak, ending before mid leaf, obscured by laminal cells almost throughout, pellucid proximally, abaxial costa cells smooth;

basal laminal cells few, pellucid, smooth, region not reaching margin;

medial cells round, 4 µm, papillae many, unbranched.

Seta

1–1.5 cm.

Capsule

oblong, urn 1.6–2.8 mm;

stomata at base;

annulus not differentiated;

operculum obliquely short-rostrate, 1.3 mm;

exostome teeth irregular, 0.3 mm, striolate proximally, trabeculate and ± finely papillose towards apex;

endostome basal membrane 3 or 4 cells high, segments well developed.

Spores

10–13 µm, densely papillose.

Perichaetia

never beyond last branching points, leaves abruptly narrowed, costa to beyond mid leaf, ending near apex in most interior leaves, laminal cells smooth.

rare, on terminal branches, leaves oblong, apex acuminate, laminal cells with 1 or 2 papillae per lumen.

Sporophytes

unknown.

Anomodon attenuatus

Anomodon tristis

Phenology Capsules mature early-mid fall.
Habitat Tree bark, at base, soil, rock Bark of trees, deciduous forests
Elevation low to moderate elevations moderate to high elevations
Distribution
from FNA
AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; MB; NB; NF; NS; ON; PE; QC; Mexico; West Indies (Cuba, Hispaniola, Jamaica); Central America (Belize, Guatemala); Europe; Asia (India, e Russia, Turkey)
[WildflowerSearch map]
from FNA
AL; AR; AZ; CT; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; VT; WI; WV; HI; NB; NS; ON; QC; Mexico (Jalisco, Nuevo León, Sonora, Veracruz); Central America (Costa Rica); South America (Bolivia); Europe; Asia
Discussion

Anomodon attenuatus is a rather polymorphic species producing abundant sporophytes in North America. The attenuate branches, sometimes arcuate and seldom ascending, are distinctive. However, this character is sometimes lacking; in some specimens the branch apices become slightly capitate and incurved. Robust plants of A. attenuatus might be mistaken for those of A. viticulosus as both have acute leaf apices. They can be distinguished by their branch morphology: prostrate and complanate in A. attenuatus, ascending and robust in A. viticulosus. Also, the leaf shape is different in both species: lanceolate and with no constriction in A. attenuatus, broadly ligulate, abruptly narrowed from a broadly ovate base in A. viticulosus. The costa in A. attenuatus also lacks the thick, aligned papillae on the dorsal surface. A consistent character in fertile plants is the absence of gametangia beyond the most recent branching points (they are never formed in the youngest branches). The branching pattern is often of several orders of successive branching, in stepwise fashion; the endostome is papillose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although the degree to which the apex breaks off is variable within Anomodon tristis, the character allows for easy identification of this species and others in sect. Haplohymenium. However, other taxa outside the section also present this feature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 634. FNA vol. 28, p. 633.
Parent taxa Anomodontaceae > Anomodon Anomodontaceae > Anomodon
Sibling taxa
A. longifolius, A. minor, A. rostratus, A. rugelii, A. thraustus, A. tristis, A. viticulosus
A. attenuatus, A. longifolius, A. minor, A. rostratus, A. rugelii, A. thraustus, A. viticulosus
Synonyms Leskea attenuata, Hypnum attenuatum Leskea tristis, Haplohymenium triste, Hypnum triste
Name authority (Hedwig) Huebener: Muscol. Germ., 562. (1833) (Cesati) Sullivant & Lesquereux: in W. S. Sullivant, Musc. Hepat. U.S., 241. (1856)
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