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anomodon moss, poodle moss

anomodon moss, yellow yarn moss

Habit Plants large, in dense mats, yellowish green. Plants small, in thick, dense, mats, light to yellowish green.
Stems

1.5–3 cm, 0.8–1 mm thick when dry, profusely branched, irregularly pinnate, primary branches prostrate to arcuate, mostly attenuate, secondary branches attenuate at apices;

central strand cells not differentiated;

pseudoparaphyllia absent;

rhizoids somewhat abundant.

0.5–2 cm, 1 mm thick when dry, profusely branched, primary branches erect;

central strand cells differentiated, smaller;

pseudoparaphyllia absent;

rhizoids few.

Branch leaves

appressed when dry, complanate when moist, ligulate, slightly narrowed mid leaf, 1.2–2(–2.2) mm;

base broadly decurrent;

margins plane, entire from base to apex (cells moderately mammillose at insertion), sometimes denticulate near apex;

apex acute, sometimes obtuse or slightly apiculate, intact;

costa strong, sharply ending a few cells before apex, sometimes fading in youngest leaves, not obscured by laminal cells distally, pellucid, abaxial costa cells smooth;

basal laminal cells hyaline or sometimes chlorophyllose, papillae few, region sometimes extending more than 1/2 length of leaf base;

distal cells quadrate, 6–8 µm, papillae many, branched.

julaceous, imbricate when dry, erect when moist, long-lanceolate, 0.8–1.4 mm;

base broadly decurrent;

margins revolute, entire;

apex acuminate, ending in hair-point, intact;

costa strong, ending well before apex, slightly obscured by laminal cells distally, pellucid, abaxial costa cells papillose, papillae thick, in 1 row;

basal laminal cells hyaline, smooth, region in basal 1/2–2/3 of lamina;

medial cells rhomboidal, 8–10(–12) µm, papillae many, slightly branched.

Seta

1–1.5 cm.

0.7–1.3 cm.

Capsule

oblong, urn 1.6–2.8 mm;

stomata at base;

annulus not differentiated;

operculum obliquely short-rostrate, 1.3 mm;

exostome teeth irregular, 0.3 mm, striolate proximally, trabeculate and ± finely papillose towards apex;

endostome basal membrane 3 or 4 cells high, segments well developed.

elliptic, ovoid to oval-cylindric, urn 1–1.5 mm;

stomata at base;

annulus of 2 rows of cells;

operculum obliquely rostrate, 0.8–1.1 mm;

peristome well developed;

exostome teeth regular, 0.2–0.3 mm, trabeculate, striolate to mid tooth or beyond, conspicuously papillose at apex;

endostome well developed, basal membrane 7+ cells high, segments long, slender.

Spores

10–13 µm, densely papillose.

11–15 µm, densely papillose.

Perichaetia

never beyond last branching points, leaves abruptly narrowed, costa to beyond mid leaf, ending near apex in most interior leaves, laminal cells smooth.

distal to last branching nodes, leaves elongate, apex long-acuminate, laminal cells smooth.

Anomodon attenuatus

Anomodon rostratus

Phenology Capsules mature early-mid fall. Capsules mature early-mid fall.
Habitat Tree bark, at base, soil, rock Base or bark of deciduous trees, soil, fallen logs, deciduous forests, open spaces, secondary forests, vertical calcareous rock, occasionally acidic
Elevation low to moderate elevations low to high elevations
Distribution
from FNA
AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; MB; NB; NF; NS; ON; PE; QC; Mexico; West Indies (Cuba, Hispaniola, Jamaica); Central America (Belize, Guatemala); Europe; Asia (India, e Russia, Turkey)
[WildflowerSearch map]
from FNA
AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; AB; BC; NB; NF; NS; ON; QC; Mexico (Chiapas, Hidalgo, Nuevo León, Puebla, San Luis Potosí); West Indies (Hispaniola, Jamaica); Bermuda; Central America (Guatemala, Honduras); Europe
[WildflowerSearch map]
Discussion

Anomodon attenuatus is a rather polymorphic species producing abundant sporophytes in North America. The attenuate branches, sometimes arcuate and seldom ascending, are distinctive. However, this character is sometimes lacking; in some specimens the branch apices become slightly capitate and incurved. Robust plants of A. attenuatus might be mistaken for those of A. viticulosus as both have acute leaf apices. They can be distinguished by their branch morphology: prostrate and complanate in A. attenuatus, ascending and robust in A. viticulosus. Also, the leaf shape is different in both species: lanceolate and with no constriction in A. attenuatus, broadly ligulate, abruptly narrowed from a broadly ovate base in A. viticulosus. The costa in A. attenuatus also lacks the thick, aligned papillae on the dorsal surface. A consistent character in fertile plants is the absence of gametangia beyond the most recent branching points (they are never formed in the youngest branches). The branching pattern is often of several orders of successive branching, in stepwise fashion; the endostome is papillose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Anomodon rostratus forms dense and thick mats, often very extensive, with clustered stems in a pseudoverticillate branching pattern. This species is distinguishable from congeners in the short branch leaves, ending in a hyaline hair-point several cells long (to 0.2+ mm) and one cell thick. The leaf areolation is also more lax; the basal portion of rhomboid cells extends beyond 1/2 the leaf length. The species fruits frequently and abundantly in North America. Despite this, male gametophytes in North America are extremely scarce, even in populations fruiting profusely. The paucity of male plants is probably the reason why sporophytes are virtually unknown elsewhere.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 634. FNA vol. 28, p. 633.
Parent taxa Anomodontaceae > Anomodon Anomodontaceae > Anomodon
Sibling taxa
A. longifolius, A. minor, A. rostratus, A. rugelii, A. thraustus, A. tristis, A. viticulosus
A. attenuatus, A. longifolius, A. minor, A. rugelii, A. thraustus, A. tristis, A. viticulosus
Synonyms Leskea attenuata, Hypnum attenuatum Leskea rostrata, Hypnum rostratum
Name authority (Hedwig) Huebener: Muscol. Germ., 562. (1833) (Hedwig) Schimper: Syn. Musc. Eur., 488. (1860)
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