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narcissus anemone, narcissus-flower anemone

Piper's anemone, Piper's windflower

Aerial shoots

7-60 cm, from caudices, caudices ascending to vertical.

10-35 cm, from rhizomes, rhizomes primarily vertical, occasionally strongly ascending.

Basal leaves

3-10, ternate;

petiole (2-)4-20 cm;

terminal leaflet ±sessile, obtriangular to oblanceolate, (2.5-)3-6(-9) × 2-10 cm, base narrowly cuneate to cuneate, margins incised (sometimes with few serrate teeth) on distal 1/3, apex acute to obtuse, surfaces glabrous or puberulous to villous or pilose;

lateral leaflets 1-3x-parted and -lobed; ultimate lobes 3-10 mm wide.

(0-)1-2, ternate;

petiole 10-20 cm;

terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2.5-6 × (1-)2-4 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous;

lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes 10-19 mm wide.

Inflorescences

2-8-flowered umbels or flowers solitary;

peduncle puberulous to villous or pilose to nearly glabrous;

involucral bracts (2-)3, 1-tiered, simple, greatly reduced, otherwise similar to basal leaves, obtriangular, distally 3-cleft and pinnatifid, (1-)1.5-5(-5.5) cm, bases clasping, ±connate, margins incised on distal 1/3, apex acuminate-acute to obtuse, surfaces glabrous or puberulous to villous or pilose;

segments primarily 3, subulate or narrowly obtriangular;

lateral segments unlobed or 2-3x-parted and -lobed; ultimate lobes 3-10 mm wide.

1-flowered;

peduncle coarsely pilose distally;

involucral bracts 3, 1-tiered, ternate, ±similar to basal leaves, bases distinct;

terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2-5.5(-7) × (0.6-)1-2.5 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous;

lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes (5-)8-18 mm wide.

Flowers

sepals 5-9, white or yellow, or abaxially white, tinged blue, white, or blue, and adaxially white, ovate to rhombic or obovate, 8-20 × 5-13(-15) mm, glabrous;

stamens 40-80(-100).

sepals 5-7, white, rarely pinkish, elliptic-obovate to ovate, (6-)8-20 × 6-8 mm, glabrous;

stamens 35-55(-90).

Achenes

body ellipsoid to ovate, flat, 5-9 × (3-)4-6 mm, winged, glabrous;

beak curved to recurved, 0.8-1.5 mm, glabrous.

body ellipsoid to obliquely ovoid, 3-4 × 1.5-2 mm, not winged, villous;

beak straight or slightly curved, 0.5-1 mm, glabrous or proximally minutely puberulous, not plumose.

Heads of achenes

spheric;

pedicel (4.5-)5-14(-18.5) cm.

nearly spheric;

pedicel (1.5-)2-5 cm.

2n

=14.

Anemone narcissiflora

Anemone piperi

Phenology Flowering spring–summer (Apr–Aug).
Habitat Shaded, moist woods
Elevation 400-3000 m (1300-9800 ft)
Distribution
from FNA
AK; CO; WY; BC; NT; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; MT; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties ca. 12 (3 in the flora).

J. Jalas (1988), W. Greuter (1989), W. Greuter et al. (1989), J. Jalas and J. Suominen (1989), and T. G. Tutin et al. (1993+, vol. 1) have recently used the name Anemone narcissifolia Linnaeus because they considered Anemone narcissiflora an illegitimate name. B. E. Dutton et al. (1995) recently proposed to conserve the orthography of Anemone narcissiflora, and the authors of this treatment ollow 14A.1 of the Code, which recommends following "existing usage as far as possible pending the General Committee's recommendation on the proposal" (W. Greuter et al. 1994).

The taxonomy of this highly variable, widespread species is extremely controversial. The conservative approach taken here most closely approximates S.L. Welsh's (1974) treatment for the Alaskan varieties. E. Hultén's discussion (1941-1950, vol. 4, pp. 735-736) of local races and the variation within this species, however, clearly illustrates the need for a thorough biosystematic investigation. Recognition of about 12 varieties is in light of S. V. Juzepczuk's (1970) work; however, he elevated local races to specific rank in his treatment.

The Aleuts used Anemone narcissiflora (no varieties specified) medicinally as an antihemorrhagic (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Anemone piperi from southeastern Washington and northeastern Oregon (i.e., the westernmost limits of the species) are sometimes intermediate between A. piperi and A. oregana. Although they possess vertical rhizomes characteristic of A. piperi, they have the bluish or pinkish sepals of A. oregana. These plants are best referred to A. piperi, pending detailed biosystematic analysis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lateral segments of involucral bracts lobed; petiole of basal leaves 15–20 cm; inflorescences umbels; stamens 60–80(–100).
var. villosissima
1. Lateral segments of involucral bracts unlobed; petiole of basal leaves (2–)4–10 cm; inflorescences umbels or flowers solitary; stamens 40–60.
→ 2
2. Sepals (when fresh) white or abaxially white or blue and adaxially white (drying yellow); body of achenes 6–9 mm.
var. monantha
2. Sepals (when fresh) yellow; body of achenes ca. 5 mm.
var. zephyra
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
Subordinate taxa
A. narcissiflora var. monantha, A. narcissiflora var. villosissima, A. narcissiflora var. zephyra
Name authority Linnaeus: Sp. Pl. 1: 542. (1753) Britton ex Rydberg: Bull. Torrey Bot. Club 29: 153. (1902)
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