Anemone multifida |
Anemone piperi |
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anémone multifide, cliff anemone, cut-leaf anemone, cut-leaf windflower, globe anemone, Pacific anemone |
Piper's anemone, Piper's windflower |
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Aerial shoots | 10-70 cm, from caudices, caudices ascending to vertical. |
10-35 cm, from rhizomes, rhizomes primarily vertical, occasionally strongly ascending. |
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Basal leaves | 3-6(-10), 1-2-ternate; petiole (2-)4-10(-14) cm; terminal leaflet petiolulate to ±sessile, broadly and irregularly rhombic to obovate, (1.5-)2.5-4.5(-5.5) × (1-)3-10 cm, base narrowly cuneate, margins incised on distal 1/3, apex broadly acute to nearly obtuse; surfaces abaxially villous-silky, hispid to villous, or sparsely long-pilose, adaxially glabrous, nearly glabrous, villous-silky, or hispid to villous; lateral leaflets (2-)3x-parted; ultimate segments (1.5-)2-3.5(-5) mm wide. |
(0-)1-2, ternate; petiole 10-20 cm; terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2.5-6 × (1-)2-4 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous; lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes 10-19 mm wide. |
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Inflorescences | 2-7-flowered cymes or flowers solitary; peduncle villous, pilose, or hispid to villous; involucral bracts usually 3-5, occasionally 2 in secondary involucres, 1-2-tiered, ternate, occasionally incompletely ternate, ±similar to basal leaves, greatly reduced, bases distinct; terminal leaflet petiolulate to ±sessile, broadly and irregularly rhombic to obovate, (1.5-)3-4(-5) × 0.5-1(-2) cm, base narrowly cuneate, margins incised on distal 1/3, apex broadly acute to nearly obtuse, surfaces abaxially hispid to villous, villous-silky, or sparsely long-pilose, adaxially glabrous, nearly glabrous, hispid to villous, or villous-silky; lateral leaflets (2-)3x-parted or -lobed, lobes frequently unequal; ultimate lobes 1.5-3(-4.3) mm wide. |
1-flowered; peduncle coarsely pilose distally; involucral bracts 3, 1-tiered, ternate, ±similar to basal leaves, bases distinct; terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2-5.5(-7) × (0.6-)1-2.5 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous; lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes (5-)8-18 mm wide. |
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Flowers | sepals 5-9, green to yellow, blue, purple, red, or occasionally white, or abaxially blue, red, yellow and red, or purple, or tinged purple, adaxially white, yellow, yellow and red, blue, or tinged purple, ovate to oblong, 5-17 × (3.5-)5-7(-9) mm, abaxially hairy, adaxially glabrous; stamens 50-80. |
sepals 5-7, white, rarely pinkish, elliptic-obovate to ovate, (6-)8-20 × 6-8 mm, glabrous; stamens 35-55(-90). |
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Achenes | body irregularly ellipsoid or elliptic, flat, 3-4 × 1.5-2 mm, not winged, tomentose, woolly, or villous; beak ±straight, distally recurved or strongly hooked, 1-6 mm, glabrous. |
body ellipsoid to obliquely ovoid, 3-4 × 1.5-2 mm, not winged, villous; beak straight or slightly curved, 0.5-1 mm, glabrous or proximally minutely puberulous, not plumose. |
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Heads of achenes | spheric; pedicel 6-15(-23) cm. |
nearly spheric; pedicel (1.5-)2-5 cm. |
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Anemone multifida |
Anemone piperi |
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Phenology | Flowering spring–summer (Apr–Aug). | |||||||||||||||||||||||||||||
Habitat | Shaded, moist woods | |||||||||||||||||||||||||||||
Elevation | 400-3000 m (1300-9800 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; South America
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ID; MT; OR; UT; WA; BC
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Discussion | Varieties 4 (4 in the flora). G.Boraiah and M.Heimburger (1964) conducted an extensive cytotaxonomic analysis of this wide-ranging and extremely variable species and its relatives. The present treatment takes a broader view of the species (and its variation) and recognizes fewer entities. In addition, Anemone tetonensis and A. stylosa, plants treated as closely related species by G.Boraiah and M.Heimburger, are treated here as varieties of A. multifida. Early-season plants of Anemone multifida var. multifida have solitary flowers and will key to var. saxicola. Anemone multifida var. tetonensis and especially var. saxicola might be based on characteristics that are influenced primarily by environment; further study is warranted. Native Americans used Anemone multifida (no varieties specified) medicinally as an antirheumatic, cold remedy, nosebleed cure, and general panacea, as well as a means of killing lice and fleas (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants of Anemone piperi from southeastern Washington and northeastern Oregon (i.e., the westernmost limits of the species) are sometimes intermediate between A. piperi and A. oregana. Although they possess vertical rhizomes characteristic of A. piperi, they have the bluish or pinkish sepals of A. oregana. These plants are best referred to A. piperi, pending detailed biosystematic analysis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3. | ||||||||||||||||||||||||||||
Parent taxa | Ranunculaceae > Anemone | Ranunculaceae > Anemone | ||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||
Name authority | Poiret: in J. Lamarck et al., in J.Lamarck et al., Encycl., suppl. 1: 364. (1810) | Britton ex Rydberg: Bull. Torrey Bot. Club 29: 153. (1902) | ||||||||||||||||||||||||||||
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