Anemone multifida |
Anemone caroliniana |
|||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
anémone multifide, cliff anemone, cut-leaf anemone, cut-leaf windflower, globe anemone, Pacific anemone |
Carolina anemone |
|||||||||||||||||||||||||||||
Aerial shoots | 10-70 cm, from caudices, caudices ascending to vertical. |
5-35(-60) cm, from tubers with rhizomes near apex, tubers ascending or vertical, rhizomes horizontal or ascending. |
||||||||||||||||||||||||||||
Basal leaves | 3-6(-10), 1-2-ternate; petiole (2-)4-10(-14) cm; terminal leaflet petiolulate to ±sessile, broadly and irregularly rhombic to obovate, (1.5-)2.5-4.5(-5.5) × (1-)3-10 cm, base narrowly cuneate, margins incised on distal 1/3, apex broadly acute to nearly obtuse; surfaces abaxially villous-silky, hispid to villous, or sparsely long-pilose, adaxially glabrous, nearly glabrous, villous-silky, or hispid to villous; lateral leaflets (2-)3x-parted; ultimate segments (1.5-)2-3.5(-5) mm wide. |
1-3(-5), 1-2-ternate; petiole 3-9(-10) cm; terminal leaflet petiolulate, rarely sessile, obtriangular, 2-3-cleft, 1-3 × 0.8-2(-2.5) cm, base narrowly attenuate to strongly and narrowly cuneate, margins serrate, rarely dissected on distal 2/3, apex obtuse to broadly acute, surfaces glabrous; lateral leaflets ±pinnately 1-2(-3)×-lobed or -parted; ultimate lobes 2-10 mm wide. |
||||||||||||||||||||||||||||
Inflorescences | 2-7-flowered cymes or flowers solitary; peduncle villous, pilose, or hispid to villous; involucral bracts usually 3-5, occasionally 2 in secondary involucres, 1-2-tiered, ternate, occasionally incompletely ternate, ±similar to basal leaves, greatly reduced, bases distinct; terminal leaflet petiolulate to ±sessile, broadly and irregularly rhombic to obovate, (1.5-)3-4(-5) × 0.5-1(-2) cm, base narrowly cuneate, margins incised on distal 1/3, apex broadly acute to nearly obtuse, surfaces abaxially hispid to villous, villous-silky, or sparsely long-pilose, adaxially glabrous, nearly glabrous, hispid to villous, or villous-silky; lateral leaflets (2-)3x-parted or -lobed, lobes frequently unequal; ultimate lobes 1.5-3(-4.3) mm wide. |
1-flowered; peduncle proximally glabrous, distally villous; involucral bracts 3, borne below middle of scape at anthesis, 1-tiered, simple, ±similar to basal leaves, obtriangular, 3-cleft, 1-2.5 cm, bases somewhat clasping, connate, margins incised on distal 1/2, apex acuminate, surfaces sparsely pilose; segments 3, broadly linear; lateral segments 1-2x-lobed or -parted; ultimate lobes 1-2.5 mm wide. |
||||||||||||||||||||||||||||
Flowers | sepals 5-9, green to yellow, blue, purple, red, or occasionally white, or abaxially blue, red, yellow and red, or purple, or tinged purple, adaxially white, yellow, yellow and red, blue, or tinged purple, ovate to oblong, 5-17 × (3.5-)5-7(-9) mm, abaxially hairy, adaxially glabrous; stamens 50-80. |
sepals 10-20(-30), white to rose or blue to purple, linear to oblong, (6-)10-20(-22) × 2-5(-7) mm, abaxially sparsely hairy, adaxially glabrous or nearly so; stamens 50-60. |
||||||||||||||||||||||||||||
Achenes | body irregularly ellipsoid or elliptic, flat, 3-4 × 1.5-2 mm, not winged, tomentose, woolly, or villous; beak ±straight, distally recurved or strongly hooked, 1-6 mm, glabrous. |
body ovoid, 1.5-2.5(-3) × ca. 2 mm, not winged, densely woolly; beak straight, 1.3-2 mm, projecting beyond achene indument, proximally tomentose, not plumose. |
||||||||||||||||||||||||||||
Heads of achenes | spheric; pedicel 6-15(-23) cm. |
fusiform; pedicel (5-)10-20 cm. |
||||||||||||||||||||||||||||
2n | =16. |
|||||||||||||||||||||||||||||
Anemone multifida |
Anemone caroliniana |
|||||||||||||||||||||||||||||
Phenology | Flowering winter–spring (Feb–Apr). | |||||||||||||||||||||||||||||
Habitat | Dry prairies, barrens, pastures, meadows, rocky, open woods | |||||||||||||||||||||||||||||
Elevation | 60-700m (200-2300ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; South America
|
AL; AR; GA; IA; IL; KS; LA; MN; MO; MS; NE; OK; SC; SD; TN; TX; WI
|
||||||||||||||||||||||||||||
Discussion | Varieties 4 (4 in the flora). G.Boraiah and M.Heimburger (1964) conducted an extensive cytotaxonomic analysis of this wide-ranging and extremely variable species and its relatives. The present treatment takes a broader view of the species (and its variation) and recognizes fewer entities. In addition, Anemone tetonensis and A. stylosa, plants treated as closely related species by G.Boraiah and M.Heimburger, are treated here as varieties of A. multifida. Early-season plants of Anemone multifida var. multifida have solitary flowers and will key to var. saxicola. Anemone multifida var. tetonensis and especially var. saxicola might be based on characteristics that are influenced primarily by environment; further study is warranted. Native Americans used Anemone multifida (no varieties specified) medicinally as an antirheumatic, cold remedy, nosebleed cure, and general panacea, as well as a means of killing lice and fleas (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Anemone caroliniana was recorded from Indiana (Vigo County); it has since been extirpated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||||||
Source | FNA vol. 3. | FNA vol. 3. | ||||||||||||||||||||||||||||
Parent taxa | Ranunculaceae > Anemone | Ranunculaceae > Anemone | ||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||
Name authority | Poiret: in J. Lamarck et al., in J.Lamarck et al., Encycl., suppl. 1: 364. (1810) | Walter: Fl. Carol., 157. (1788) | ||||||||||||||||||||||||||||
Web links |
|