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alpine anemone, Drummond's anemone, Drummond's windflower

anémone de virginie, riverbank anemone, tall anemone, tall thimbleweed, tall windflower, thimbleweed, Virginia anemone

Aerial shoots

(7-)10-25(-30) cm, from caudices, caudices ascending to primarily vertical.

30-100(-110) cm, from caudices, rarely with ascending rhizomes, caudices ascending to vertical.

Basal leaves

5-15, 2-ternate, occasionally irregularly so;

petiole 2-10 cm;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to obtriangular, 0.5-3 × 0.5-2 cm, base narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.6 mm wide.

1-5, ternate;

petiole 5-35 cm;

terminal leaflet sessile or nearly so, oblanceolate to obovate, 2-9(-12) × 2-5(-7) cm, base cuneate to broadly cuneate, margins coarsely serrate and incised on distal 1/2, apex acuminate to narrowly acute, surfaces pilose, more so abaxially;

lateral leaflets usually 1-2x-lobed or -parted, occasionally unlobed; ultimate lobes 10-30(-40) mm wide.

Inflorescences

1(-2)-flowered;

peduncle villous;

involucral bracts 3(-4), 1-tiered, ±similar to basal leaves, highly reduced, 2-ternate or irregularly so, bases distinct;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to pinnatifid, 1-3.5 × 0.5-2 cm, bases narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.5 mm wide.

(1-)3-9-flowered cymes;

peduncle villous;

primary involucral bracts 3(-5), secondary involucral bracts 2(-3), (1-)2-tiered, ternate, ±similar to basal leaves, bases distinct;

terminal leaflet ±sessile, elliptic to oblanceolate, 2-10(-12) cm (2 cm in secondary involucre) × 2-5(-7) cm, bases cuneate, margins coarsely serrate and incised on distal 1/2, apex acuminate to narrowly acute, surfaces pilose, more so abaxially;

lateral leaflets unlobed or 1x-lobed or -parted; ultimate lobes 8-25(-35)mm wide.

Flowers

sepals (5-)6-9, white, or abaxially white, tinged blue, and adaxially white, ovate, rarely oblong or narrowly obovate, 8-20 × 6-10 mm, abaxially hairy, rarely glabrous, adaxially glabrous;

stamens 80-100, whitish;

styles white.

sepals usually 5, green, yellow, or red (rarely white or abaxially green to green-yellow and adaxially green or yellow and tinged red), oblong, ovate, or obovate, 6.5-20 × 2.5-10mm, abaxially hairy, adaxially glabrous or nearly so;

stamens 50-70.

Achenes

body ovoid, 2-4 × 1-1.5 mm, not winged, woolly;

beak straight, 2-4(-6) mm, glabrous.

body obovoid, 2-3.7 × 1.5-2mm, not winged, densely woolly;

beak curved, 1-1.5mm, puberulous, not plumose.

Heads of achenes

spheric, rarely cylindric;

pedicel (2-)3-10 cm.

oblong-ellipsoid, rarely obconic;

pedicel 13-25(-30) cm.

Anemone drummondii

Anemone virginiana

Distribution
from FNA
AK; CA; ID; MT; OR; WA; WY; AB; BC; NT; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; VA; VT; WI; WV; WY; AB; BC; MB; NB; NF; NS; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Anemone drummondii is an extremely variable species whose circumscription is controversial. Some plants appear intermediate between this species and A. multifida; cytologically the two are quite distinct (G.Boraiah and M.Heimburger 1964; C. L. Hitchcock et al. 1955-1969, vol.2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

See C. S. Keener et al. (1995) for an analysis of infraspecific variation within Anemone virginiana from which the current treatment has been adopted.

Varieties of Anemone virginiana used medicinally by native Americans were not specified; the species was used as an antidiarrheal, an aid for whooping cough, a stimulant, an emetic, a love potion, a remedy for tuberculosis, and a protection against witchcraft medicine (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ultimate segments of lateral leaflets 1–1.5(–2) mm wide, leaflets villous.
var. drummondii
1. Ultimate segments of lateral leaflets 1.5–2.6 mm wide, leaflets nearly glabrous or pilose.
var. lithophila
1. Sepals usually 5–10mm, abaxially densely tomentose; anthers 0.7–1(–1.2) mm; primarily Canadian, in dry woods, sandy ridges, and grasslands.
var. cylindroidea
1. Sepals (6–)10–21 mm, abaxially usually thinly pubescent; anthers (0.8–)1–1.7 mm; widely distributed, in moist habitats.
→ 2
2. Base of involucral bracts cordate or reniform, rarely subtruncate, terminal leaflets light green, margins proximally mostly straight- to convex-sided, variously lobed or serrate, variously pubescent; anthers typically greater than 1.1 mm; heads of achenes ovoid to ovoid-cylindric, (9–)11–14 mm diam.; widely distributed.
var. virginiana
2. Base of involucral bracts usually truncate to subtruncate, sometimes reniform or cordate, terminal leaflets deep green, margins proximally concave- to straight-sided, distally incised, thinly pubescent; anthers typically less than 1.2 mm; heads of achenes ±ovoid-cylindric, 8–10(–11) mm diam.; distributed primarily in New England, Great Lakes area, and adjacent Canada.
var. alba
Source FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford. FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa
Subordinate taxa
A. drummondii var. drummondii, A. drummondii var. lithophila
A. virginiana var. alba, A. virginiana var. cylindroidea, A. virginiana var. virginiana
Name authority S. Watson: Bot. California 2: 424. (1880) Linnaeus: Sp. Pl. 1: 540. (1753)
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