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alpine anemone, Drummond's anemone, Drummond's windflower

desert anemone, desert windflower, tuber anemone

Aerial shoots

(7-)10-25(-30) cm, from caudices, caudices ascending to primarily vertical.

10-30(-40) cm, from caudex-like tubers, tubers ascending to vertical.

Basal leaves

5-15, 2-ternate, occasionally irregularly so;

petiole 2-10 cm;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to obtriangular, 0.5-3 × 0.5-2 cm, base narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.6 mm wide.

1-3(-5), 1-2-ternate;

petiole 5-7 cm;

terminal leaflet sessile, rarely petiolulate, irregularly oblanceolate, (1.5-)2-3(-3.5) × 1-2(-2.5) cm, base narrowly cuneate, margins incised to dissected on distal 2/3, apex broadly acute, surfaces nearly glabrous;

lateral leaflets 1-2x-parted and/or -lobed; ultimate lobes 4-8(-12) mm wide.

Inflorescences

1(-2)-flowered;

peduncle villous;

involucral bracts 3(-4), 1-tiered, ±similar to basal leaves, highly reduced, 2-ternate or irregularly so, bases distinct;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to pinnatifid, 1-3.5 × 0.5-2 cm, bases narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.5 mm wide.

2-3(-5)-flowered cymes or flowers solitary;

peduncle proximally nearly glabrous, distally villous;

involucral bracts primarily 3, (1-)2-tiered, simple, ±similar to basal leaves, pinnatifid, 1.5-5.5 cm (primary involucral bracts 2-5.5 cm, secondary involucral bracts 1.5-3.5 cm), bases distinct, broadly to narrowly cuneate to clasping, margins irregularly serrulate and pinnatifid on ca. distal 1/2, apex narrowly acute to acuminate, surfaces thinly pilose;

segments primarily 3, linear to pinnatifid;

lateral segments 1-2x-parted and/or -lobed; ultimate lobes 1.5-2.5 mm wide.

Flowers

sepals (5-)6-9, white, or abaxially white, tinged blue, and adaxially white, ovate, rarely oblong or narrowly obovate, 8-20 × 6-10 mm, abaxially hairy, rarely glabrous, adaxially glabrous;

stamens 80-100, whitish;

styles white.

sepals 8-10, pink to white, linear-oblong, 10-14(-20) × (2-)3-5(-6) mm, sparsely hairy;

stamens 50-60.

Achenes

body ovoid, 2-4 × 1-1.5 mm, not winged, woolly;

beak straight, 2-4(-6) mm, glabrous.

body orbiculate, flat, 2.5-3.5 × 2-2.5 mm, not winged, densely villous;

beak straight, ca. 1.5 mm, minutely puberulous, not plumose.

Heads of achenes

spheric, rarely cylindric;

pedicel (2-)3-10 cm.

fusiform;

pedicel (5-)7-15(-22) cm.

2n

=16.

Anemone drummondii

Anemone tuberosa

Phenology Flowering spring (Apr–May).
Habitat Rocky slopes, streamsides
Elevation 800-2500 m [2600-8200 ft]
Distribution
from FNA
AK; CA; ID; MT; OR; WA; WY; AB; BC; NT; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; TX; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Anemone drummondii is an extremely variable species whose circumscription is controversial. Some plants appear intermediate between this species and A. multifida; cytologically the two are quite distinct (G.Boraiah and M.Heimburger 1964; C. L. Hitchcock et al. 1955-1969, vol.2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The tuberous anemones in North America (A. tuberosa, A. edwardsiana, A. okennonii, A. berlandieri, and A. caroliniana) are closely related to each other and to the South American species A. decapetala Arduino, A. triternata Vahl, A. cicutifolia I.M. Johnston, and A. sphenophylla Poeppig. Particularly useful characters in identifying North American plants are number of tiers of involucral bracts, presence or absence of heterophylly between basal leaves and involucral bracts, and presence or absence of rhizomes. The current treatment of this group primarily follows C. S. Keener and B. E. Dutton (1994), who included a discussion of the relationships among its members. For a cytotaxonomic discussion of most members of this group, see C. Joseph and M. Heimburger (1966).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ultimate segments of lateral leaflets 1–1.5(–2) mm wide, leaflets villous.
var. drummondii
1. Ultimate segments of lateral leaflets 1.5–2.6 mm wide, leaflets nearly glabrous or pilose.
var. lithophila
Source FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford. FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. virginiana
Subordinate taxa
A. drummondii var. drummondii, A. drummondii var. lithophila
Name authority S. Watson: Bot. California 2: 424. (1880) Rydberg: Bull. Torrey Bot. Club 29: 151. (1902)
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