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alpine anemone, Drummond's anemone, Drummond's windflower

Piper's anemone, Piper's windflower

Aerial shoots

(7-)10-25(-30) cm, from caudices, caudices ascending to primarily vertical.

10-35 cm, from rhizomes, rhizomes primarily vertical, occasionally strongly ascending.

Basal leaves

5-15, 2-ternate, occasionally irregularly so;

petiole 2-10 cm;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to obtriangular, 0.5-3 × 0.5-2 cm, base narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.6 mm wide.

(0-)1-2, ternate;

petiole 10-20 cm;

terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2.5-6 × (1-)2-4 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous;

lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes 10-19 mm wide.

Inflorescences

1(-2)-flowered;

peduncle villous;

involucral bracts 3(-4), 1-tiered, ±similar to basal leaves, highly reduced, 2-ternate or irregularly so, bases distinct;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to pinnatifid, 1-3.5 × 0.5-2 cm, bases narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.5 mm wide.

1-flowered;

peduncle coarsely pilose distally;

involucral bracts 3, 1-tiered, ternate, ±similar to basal leaves, bases distinct;

terminal leaflet sessile, rhombic, lanceolate, or oblanceolate, (1.5-)2-5.5(-7) × (0.6-)1-2.5 cm, base narrowly cuneate, margins coarsely serrate to coarsely dentate on distal 1/2-2/3, apex acuminate to narrowly acute, surfaces pilose or glabrous;

lateral leaflets unlobed or sometimes 1x-lobed; ultimate lobes (5-)8-18 mm wide.

Flowers

sepals (5-)6-9, white, or abaxially white, tinged blue, and adaxially white, ovate, rarely oblong or narrowly obovate, 8-20 × 6-10 mm, abaxially hairy, rarely glabrous, adaxially glabrous;

stamens 80-100, whitish;

styles white.

sepals 5-7, white, rarely pinkish, elliptic-obovate to ovate, (6-)8-20 × 6-8 mm, glabrous;

stamens 35-55(-90).

Achenes

body ovoid, 2-4 × 1-1.5 mm, not winged, woolly;

beak straight, 2-4(-6) mm, glabrous.

body ellipsoid to obliquely ovoid, 3-4 × 1.5-2 mm, not winged, villous;

beak straight or slightly curved, 0.5-1 mm, glabrous or proximally minutely puberulous, not plumose.

Heads of achenes

spheric, rarely cylindric;

pedicel (2-)3-10 cm.

nearly spheric;

pedicel (1.5-)2-5 cm.

Anemone drummondii

Anemone piperi

Phenology Flowering spring–summer (Apr–Aug).
Habitat Shaded, moist woods
Elevation 400-3000 m [1300-9800 ft]
Distribution
from FNA
AK; CA; ID; MT; OR; WA; WY; AB; BC; NT; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; MT; OR; UT; WA; BC
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Anemone drummondii is an extremely variable species whose circumscription is controversial. Some plants appear intermediate between this species and A. multifida; cytologically the two are quite distinct (G.Boraiah and M.Heimburger 1964; C. L. Hitchcock et al. 1955-1969, vol.2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Anemone piperi from southeastern Washington and northeastern Oregon (i.e., the westernmost limits of the species) are sometimes intermediate between A. piperi and A. oregana. Although they possess vertical rhizomes characteristic of A. piperi, they have the bluish or pinkish sepals of A. oregana. These plants are best referred to A. piperi, pending detailed biosystematic analysis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ultimate segments of lateral leaflets 1–1.5(–2) mm wide, leaflets villous.
var. drummondii
1. Ultimate segments of lateral leaflets 1.5–2.6 mm wide, leaflets nearly glabrous or pilose.
var. lithophila
Source FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford. FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
Subordinate taxa
A. drummondii var. drummondii, A. drummondii var. lithophila
Name authority S. Watson: Bot. California 2: 424. (1880) Britton ex Rydberg: Bull. Torrey Bot. Club 29: 153. (1902)
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