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alpine anemone, Drummond's anemone, Drummond's windflower

Okennon's anemone

Aerial shoots

(7-)10-25(-30) cm, from caudices, caudices ascending to primarily vertical.

20-30 cm, from tubers, tubers ±vertical.

Basal leaves

5-15, 2-ternate, occasionally irregularly so;

petiole 2-10 cm;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to obtriangular, 0.5-3 × 0.5-2 cm, base narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.6 mm wide.

4-10, 2-3-ternate or irregularly so;

petiole 5-10 cm;

terminal leaflet petiolulate, obovate, 1.2-1.8(-20) × ca. 1.5(-2.5) cm, base narrowly cuneate, margins coarsely serrate to nearly incised on distal 1/2, ciliate or not, apex broadly acute, surfaces nearly glabrous;

lateral leaflets 1-2x-lobed or -parted; ultimate lobes 2-5 mm wide.

Inflorescences

1(-2)-flowered;

peduncle villous;

involucral bracts 3(-4), 1-tiered, ±similar to basal leaves, highly reduced, 2-ternate or irregularly so, bases distinct;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to pinnatifid, 1-3.5 × 0.5-2 cm, bases narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.5 mm wide.

(1-)2-3-flowered cymes;

peduncle proximally nearly glabrous, distally downy;

involucral bracts primarily 3, (1-)2-tiered, simple, dissimilar to basal leaves, obtriangular, 3-cleft to pinnatifid, 2-5 cm, bases clasping, ±connate, margins incised throughout, apex acuminate, surfaces appressed-pilose;

segments primarily 3, linear;

lateral segments unlobed or 1x-lobed; ultimate segments 1-2 mm wide.

Flowers

sepals (5-)6-9, white, or abaxially white, tinged blue, and adaxially white, ovate, rarely oblong or narrowly obovate, 8-20 × 6-10 mm, abaxially hairy, rarely glabrous, adaxially glabrous;

stamens 80-100, whitish;

styles white.

sepals 7-11, greenish white to abaxially reddish, oblong, 6-12(-14) × (2-)3-4.5 mm, abaxially hairy, adaxially glabrous;

stamens 35-55.

Achenes

body ovoid, 2-4 × 1-1.5 mm, not winged, woolly;

beak straight, 2-4(-6) mm, glabrous.

body ovate, flat, not winged, densely white-villous (with long tuft of hairs at base);

beak ±straight, ca. 1 mm, hidden in achene indument, white-tomentose, not plumose.

Heads of achenes

spheric, rarely cylindric;

pedicel (2-)3-10 cm.

oblong-ellipsoid;

pedicel 10-18 cm.

Anemone drummondii

Anemone okennonii

Phenology Flowering spring (Mar–Apr).
Habitat Dry, open ledges and slopes
Elevation 500-1500m [1600-4900ft]
Distribution
from FNA
AK; CA; ID; MT; OR; WA; WY; AB; BC; NT; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
Discussion

Varieties 2 (2 in the flora).

Anemone drummondii is an extremely variable species whose circumscription is controversial. Some plants appear intermediate between this species and A. multifida; cytologically the two are quite distinct (G.Boraiah and M.Heimburger 1964; C. L. Hitchcock et al. 1955-1969, vol.2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ultimate segments of lateral leaflets 1–1.5(–2) mm wide, leaflets villous.
var. drummondii
1. Ultimate segments of lateral leaflets 1.5–2.6 mm wide, leaflets nearly glabrous or pilose.
var. lithophila
Source FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford. FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
Subordinate taxa
A. drummondii var. drummondii, A. drummondii var. lithophila
Synonyms A. tuberosa var. texana
Name authority S. Watson: Bot. California 2: 424. (1880) Keener & B. E. Dutton: Sida 16: 198. (1994)
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