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alpine anemone, Drummond's anemone, Drummond's windflower

Edwards Plateau anemone, Edwards Plateau thimbleweed

Aerial shoots

(7-)10-25(-30) cm, from caudices, caudices ascending to primarily vertical.

(20-)30-50 cm, from tubers, tubers ±vertical.

Basal leaves

5-15, 2-ternate, occasionally irregularly so;

petiole 2-10 cm;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to obtriangular, 0.5-3 × 0.5-2 cm, base narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.6 mm wide.

3-6, 1-(nearly 2)-ternate;

petiole 8-15 cm;

terminal leaflet petiolulate, oblanceolate to obovate, 2-5 × 2-2.5 cm, base broadly cuneate, margins coarsely crenate on distal 1/2, apex obtuse to broadly acute, surfaces glabrous;

lateral leaflets 1x-lobed; ultimate lobes 4-8(-12) mm wide.

Inflorescences

1(-2)-flowered;

peduncle villous;

involucral bracts 3(-4), 1-tiered, ±similar to basal leaves, highly reduced, 2-ternate or irregularly so, bases distinct;

terminal leaflet sessile or basally attenuate and appearing petiolulate, obovate to pinnatifid, 1-3.5 × 0.5-2 cm, bases narrowly cuneate to cuneate, margins incised to dissected on distal 1/3-1/2, apex broadly acute to obtuse, surfaces villous;

lateral leaflets 2x-parted, division frequently irregular; ultimate segments 1-2.5 mm wide.

(1-)2-5-flowered cymes;

peduncle proximally nearly glabrous, distally densely pubescent;

involucral bracts primarily 3, 2-tiered, simple, dissimilar to basal leaves, 3-cleft to pinnatifid, 2-5 cm, bases clasping, ±connate, margins coarsely crenate throughout, apex narrowly acute to acuminate, surfaces glabrous;

segments primarily 3, linear;

lateral segments unlobed or occasionally 1x-lobed; ultimate lobes 3-5 mm wide.

Flowers

sepals (5-)6-9, white, or abaxially white, tinged blue, and adaxially white, ovate, rarely oblong or narrowly obovate, 8-20 × 6-10 mm, abaxially hairy, rarely glabrous, adaxially glabrous;

stamens 80-100, whitish;

styles white.

sepals (8-)10-20, white to bluish, oblanceolate, 10-16 × 2-3 mm, sparsely hairy to nearly glabrous;

stamens 50-60.

Achenes

body ovoid, 2-4 × 1-1.5 mm, not winged, woolly;

beak straight, 2-4(-6) mm, glabrous.

body orbiculate, flat, not winged, ±dull or varnished, woolly with long tuft of hairs at base, or glabrous;

beak straight, rarely curved, 0.5-1 mm, glabrous or sparsely villous proximally, not plumose.

Heads of achenes

spheric, rarely cylindric;

pedicel (2-)3-10 cm.

obconic to fusiform;

pedicel 10-20 cm.

2n

=16.

Anemone drummondii

Anemone edwardsiana

Distribution
from FNA
AK; CA; ID; MT; OR; WA; WY; AB; BC; NT; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Anemone drummondii is an extremely variable species whose circumscription is controversial. Some plants appear intermediate between this species and A. multifida; cytologically the two are quite distinct (G.Boraiah and M.Heimburger 1964; C. L. Hitchcock et al. 1955-1969, vol.2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Anemone edwardsiana is found only in Texas on the Edwards Plateau.

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ultimate segments of lateral leaflets 1–1.5(–2) mm wide, leaflets villous.
var. drummondii
1. Ultimate segments of lateral leaflets 1.5–2.6 mm wide, leaflets nearly glabrous or pilose.
var. lithophila
1. Achenes woolly, ± dull.
var. edwardsiana
1. Achenes glabrous, varnished.
var. petraea
Source FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford. FNA vol. 3. Treatment authors: Bryan E. Dutton, Carl S. Keener, Bruce A. Ford.
Parent taxa Ranunculaceae > Anemone Ranunculaceae > Anemone
Sibling taxa
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. edwardsiana, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
A. acutiloba, A. americana, A. berlandieri, A. canadensis, A. caroliniana, A. cylindrica, A. deltoidea, A. drummondii, A. grayi, A. lancifolia, A. lyallii, A. multiceps, A. multifida, A. narcissiflora, A. occidentalis, A. okennonii, A. oregana, A. parviflora, A. patens, A. piperi, A. quinquefolia, A. richardsonii, A. tuberosa, A. virginiana
Subordinate taxa
A. drummondii var. drummondii, A. drummondii var. lithophila
A. edwardsiana var. edwardsiana, A. edwardsiana var. petraea
Name authority S. Watson: Bot. California 2: 424. (1880) Tharp: Amer. Midl. Naturalist 33: 669. (1945)
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