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bushy beardgrass, bushy bluestem

bluestem

Habit Plants cespitose, upper portion dense, oblong to oblanceolate or obpyramidal. Plants perennial; usually cespitose, sometimes rhizomatous.
Culms

20-250 cm;

internodes green, sometimes glaucous;

branches mostly erect, straight.

20-310 cm, erect, much-branched distally.

Sheaths

usually scabrous, sometimes smooth;

ligules 0.6-2.2 mm, sometimes ciliate, cilia to 0.9 mm;

blades 13-109 cm long, 2.9-9.5 mm wide, glabrous or sparsely to densely pubescent, hairs usually spreading, rarely appressed.

Leaves

not aromatic;

ligules membranous, sometimes ciliate;

blades linear, flat, folded, or convolute.

Inflorescences/Inflorescence units

10-600 per culm; subtending sheaths (2.0)2.9-4.4(6.5) cm long, (1.5)2.3-3.4(4.4) mm wide;

peduncles (1)6-14(60) mm, with 2(4) rames;

rames (1)1.7-2.5(3.5) cm, exserted or not at maturity, pubescence sparse basally and increasing in density distally within each internode.

terminal and axillary or a false panicle;

inflorescence units 1-600+ per culm;

peduncles initially concealed by the subtending leaf sheaths, sometimes exserted beyond the sheaths at maturity, with (1)2-5(13) rames;

rames not reflexed at maturity, axes slender, terete to flattened, not longitudinally grooved, usually conspicuously pubescent, with spikelets in heterogamous sessile-pedicellate pairs (the terminal spikelets sometimes in triplets of 1 sessile and 2 pedicellate spikelets), apices of the internodes neither cupulate nor fimbriate;

disarticulation in the rames, below the sessile spikelets.

Pedicels

usually longer than 3 mm, similar to the rame internodes in shape, length, and pubescence color, not fused to the rame axes.

Sessile

spikelets 3-5 mm;

callus hairs 1-2.5 mm;

keels of lower glumes sometimes scabrous below midlength, usually scabrous distally;

awns 6-19 mm;

anthers 1(3), 0.5-1.5 mm, yellow, red, or purple.

spikelets bisexual, awned, with short, blunt calluses;

lower glumes 2-keeled, flat or concave, usually not veined between the keels, sometimes 2-9-veined;

anthers 1, 3(2).

Pedicellate

spikelets vestigial or absent, sterile.

spikelets usually vestigial or absent, sometimes well-developed and staminate.

x

= 10.

2n

= 20.

Andropogon glomeratus

Andropogon

Distribution
from FNA
AL; AR; AZ; CA; DC; DE; FL; GA; IL; KY; LA; MA; MD; MS; NC; NJ; NM; NV; NY; OH; OK; PA; RI; SC; TN; TX; UT; VA; WV; HI; PR; Virgin Islands
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from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; WY; HI; PR; MB; ON; QC; SK; Virgin Islands
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Discussion

Andropogon glomeratus hybridizes with both A. longiberbis and A. virginicus. Some of its varieties are morphologically similar to the latter species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Andropogon is a cosmopolitan genus of tropical and temperate zones, comprising approximately 120 species. Thirteen species are native to the Flora region. Andropogon bicornis has been found in the region but is not known to be established. All but A. hallii grow in the southeastern United States.

Several taxa are ecologically important in North America. Andropogon gerardii is one of the most important native grasses in North America, being one of the dominant species in the tall-grass prairies that used to cover the center of the continent. Many varieties of A. glomeratus and A. virginicus aggressively colonize abandoned fields, cutover timberlands, and roadsides. Some species are used in restoration and landscaping.

Species of Andropogon with solitary rames are easily confused with Schizachyrium but, in Andropogon, the lower glumes of the sessile spikelets are flat or concave and the rame internodes are not cupulate, whereas Schizachyrium has convex glumes and rame internodes with strongly cupulate apices. Successful identification of species in Andropogon sect. Leptopogon (numbers 3-14) requires mature, complete specimens and careful field study (Campbell 1983, 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Blades glaucous, glabrous, and smooth
var. glaucopsis
1. Blades green, often pubescent or scabrous
→ 2
2. Sheaths subtending the inflorescence units 1.5-3 mm wide; leaf sheaths usually smooth; ligules ciliate, the cilia 0.2-0.9 mm long
var. pumilus
2. Sheaths subtending the inflorescence units (1.5)2.3-3.4(4.4) mm wide; leaf sheaths often scabrous; ligules, when ciliate, with the cilia no more than 0.5 mm long.
→ 3
3. Keels of the lower glumes scabrous below and beyond midlength
var. scabriglumis
3. Keels of the lower glumes usually smooth below midlength, scabrous distally.
→ 4
4. Upper portion of the plants oblong to obpyramidal; mature peduncles (4)11-35(60) mm long; anthers eventually falling
var. glomeratus
4. Upper portion of the plants cylindrical to oblong; mature peduncles 2-5 (8) mm long; withered remnants of anthers retained within the spikelets
var. hirsutior
1. Pedicellate spikelets usually well-developed, (3.5)6-12 mm long, usually staminate; sessile spikelets 5-12 mm long (sect. Andropogon).
→ 2
2. Sessile spikelets with awns 8-25 mm long; ligules 0.4-2.5 mm long; hairs of the rame internodes 2.2-4.2 mm long, sparse to dense; rhizomes sometimes present, the internodes usually less than 2 cm
A. gerardii
2. Sessile spikelets unawned or with awns less than 11 mm long; ligules (0.9)2.5-4.5 mm long; hairs of the rame internodes 3.7-6.6 mm long, usually dense; rhizomes always present, the internodes often more than 2 cm long
A. hallii
1. Pedicellate spikelets usually vestigial or absent, those of the terminal spikelet units occasionally well-developed and staminate; sessile spikelets 2.6-8.4 mm long (sect. Leptopogon).
→ 3
3. Peduncles with solitary rames; plants of southern Florida
A. gracilis
3. Peduncles with (1)2-13 rames; plants of varied distribution, including southern Florida.
→ 4
4. Rames not or scarcely exserted at maturity; peduncles mostly less than 15 mm long at maturity.
→ 5
5. Culms 30-140 (usually about 80) cm tall; blades 0.8-5 (usually about 2.5) mm wide; inflorescence units 2-31 per culm
A. gyrans
5. Culms 20-250 (usually more than 90) cm tall; blades 1.7-9.5 (usually more than 3) mm wide; inflorescence units 3-600 per culm.
→ 6
6. Blades pubescent, most hairs appressed; callus hairs 1.5-5 mm long
A. longiberbis
6. Blades glabrous or with spreading (rarely appressed) hairs; callus hairs 1-3 mm long.
→ 7
7. Blades 11-52 cm long; sheaths smooth, rarely somewhat scabrous; ligules 0.2-1 mm long; keels of the lower glumes usually smooth below midlength, scabrous distally
A. virginicus
7. Blades 13-109 cm long; sheaths usually scabrous; ligules 0.6-2.2 mm long; keels of the lower glumes sometimes scabrous below midlength
A. glomeratus
4. Rames sometimes exserted above their subtending sheaths at maturity; 1 or more peduncles more than 15 mm long at maturity.
→ 8
8. Anthers 3.
→ 9
9. Sessile spikelets 4.5-8.4 mm long; pedicellate spikelets 1.5-3.6 mm long, sterile; plants common and widespread in the southeastern United States
A. ternarius
9. Sessile spikelets 3-4 mm long; pedicellate spikelets mostly vestigial or absent, those of the terminal spikelet units well-developed and staminate; in the Flora region, known only from southern Florida
A. bicornis
8. Anthers 1 (rarely 3).
→ 10
10. Peduncles with 2-13 rames
A. liebmannii
10. Peduncles usually with 2 (infrequently up to 4) rames or (in A. gyrans var. gyrans and A. virginicus var. virginicus), 2-5 (infrequently up to 7) rames.
→ 11
11. Culms 30-120(140) (usually less than 100) cm tall; blades 0.8-5 (usually less than 3) mm wide; inflorescence units 2-31 per culm.
→ 12
12. Peduncles with 2-5 rames; anthers 0.6-1.7 mm long; sessile spikelets (3)4.1-4.4(5.7) mm long
A. gyrans
12. Peduncleswith 2 rames; anthers 1.2-2 mm long; sessile spikelets (4)4.8-5(5.5) mm long
A. tracyi
11. Culms (20)90-310 (usually more than 100) cm tall; blades 1.7-9.5 (usually more than 3) mm wide; inflorescence units 5-210 per culm.
→ 13
13. Upper portion of the plants open, the branches conspicuously arching
A. brachystachyus
13. Upper portion of the plants dense, the branches usually straight and erect to ascending.
→ 14
14. Rame internodes usually densely and uniformly pubescent over their entire length; anthers 1.3-3.5 mm long; sessile spikelets (3.8)4-6.1 mm long.
→ 15
15. Blades 15-35 cm long, often more or less pubescent; sheaths smooth, very rarely somewhat scabrous; anthers 2-3.5 mm long; inflorescence units 5-45 per culm
A. arctatus
15. Blades 32-61 cm long, usually glabrous; sheaths often scabrous; anthers 1.3-2 mm long; inflorescence units usually at least 50 (9-210) per culm
A. floridanus
14. Rame internodes sparsely pubescent basally, more densely pubescent distally; anthers 0.5-1.5 mm long; sessile spikelets 2.6-4(5).
→ 16
16. Blades 11-52 cm long; sheaths smooth, rarely somewhat scabrous; ligules 0.2-1 mm long; keels of the lower glumes usually smooth below midlength, scabrous distally
A. virginicus
16. Blades 13-109 cm long; sheaths usually scabrous, sometimes smooth; ligules 0.6-2.2 mm long; keels of the lower glumes sometimes scabrous below midlength
A. glomeratus
Source FNA vol. 25, p. 661. FNA vol. 25, p. 649. Author: Christopher S. Campbell;.
Parent taxa Poaceae > subfam. Panicoideae > tribe Andropogoneae > Andropogon > sect. Leptopogon Poaceae > subfam. Panicoideae > tribe Andropogoneae
Sibling taxa
A. arctatus, A. bicornis, A. brachystachyus, A. floridanus, A. gerardii, A. gracilis, A. gyrans, A. hallii, A. liebmannii, A. longiberbis, A. ternarius, A. tracyi, A. virginicus
Subordinate taxa
A. glomeratus var. glaucopsis, A. glomeratus var. glomeratus, A. glomeratus var. hirsutior, A. glomeratus var. pumilus, A. glomeratus var. scabriglumis
A. arctatus, A. bicornis, A. brachystachyus, A. floridanus, A. gerardii, A. glomeratus, A. gracilis, A. gyrans, A. hallii, A. liebmannii, A. longiberbis, A. ternarius, A. tracyi, A. virginicus
Name authority (Walter) Britton, Sterns & Poggenb. L.
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