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anacolia moss

anacolia moss

Habit Plants yellowish or reddish. Plants in lax to dense tufts, sometimes viny, reddish brown proximally, yellowish green to green distally.
Stems

1–5 cm.

1–5 cm, erect to ascending, simple or fastigiate;

octagonal in cross section, hyalodermis absent, epidermis prorulose;

radiculose proximally, rhizoids papillose.

Leaves

imbricate or distal leaves divaricate when dry, spreading to recurved when moist, narrowly ovate-lanceolate to lanceolate, distal lamina 2-stratose toward costa, 3- or 4-stratose at margins, 2–6.5 mm;

margins coarsely doubly serrate distally;

costa excurrent to long-excurrent, abaxial surface rough;

inner basal laminal cells elongated oblong;

distal cells short-rectangular to linear, to 45 × 3–7 µm, prorulose at both ends.

not in distinct rows or rarely in 5 rows, imbricate, appressed when dry, erect to spreading or recurved when moist, narrowly lanceolate or ovate-lanceolate, 1–3-stratose (sometimes 3- or 4-stratose at margins);

base not sheathing;

margins revolute in distal 1/2, sometimes almost to base, serrate, teeth single or paired;

apex acute, acuminate, or rarely obtuse;

costa percurrent to long-excurrent, abaxial surface rough or smooth;

basal laminal cells shorter, wider than distal cells;

distal cells usually oblong to linear or quadrate, strongly prorulose to nearly smooth, walls thick.

Seta

single, short to elongate, straight or somewhat curved.

Sexual condition

dioicous;

perigonia gemmiform;

perichaetial leaves scarcely distinct.

Capsule

2–3 mm;

operculum short-conic;

peristome absent.

erect to inclined, globose to ovoid, irregularly wrinkled when dry, mouth small;

annulus absent or scarcely developed;

operculum short-conic to conic-obtuse;

peristome absent, of low membrane, or exostome teeth yellow to reddish brown, short, fragile, apically free;

endostome rare, very faint.

Spores

23–28 µm].

globose to reniform, warty or coarsely papillose.

[seta 0.2–0.8 cm.

Specialized

asexual reproduction unknown.

Anacolia laevisphaera

Anacolia

Habitat Dry to moist soil, rock crevices, talus slopes
Elevation moderate to high elevations (1000-1700 m) (moderate to high elevations (3300-5600 ft))
Distribution
from FNA
AZ; CO; NM; Mexico; Central America; South America; West Indies; Asia; Africa
[WildflowerSearch map]
North America; Mexico; Central America; South America; West Indies; Europe; Asia; Africa
Discussion

The prominently prorulose laminal cells and well-differentiated cells of the leaf base distinguish Anacolia laevisphaera from A. menziesii. When sterile, Bartramia stricta may be mistaken for A. laevisphaera. The basal laminal cells of this species are subquadrate, to short-rectangular toward the margins, and 20 × 10–12 µm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 4 (2 in the flora).

Anacolia is recognized by its imbricate, appressed leaves, borne on prorulose stems that are felty tomentose proximally. The basal juxtacostal laminal cells are sometimes linear; the perigonia and perichaetia are terminal, appearing lateral by innovations; the capsule is leptodermous; and the capsule neck is very short or absent. When fertile, the usually erect, irregularly wrinkled capsules are distinctive. Within the flora area, Anacolia is confined to western North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal laminal cells prorulose at both ends on both surfaces; inner basal cells elongated oblong; distal lamina 2-4-stratose.
A. laevisphaera
1. Distal laminal cells smooth or a few cells bearing low prorulae on abaxial side; inner basal cells quadrate or short-rectangular; distal lamina 1 (or 2)-stratose.
A. menziesii
Source FNA vol. 28, p. 101. FNA vol. 28, p. 99.
Parent taxa Bartramiaceae > Anacolia Bartramiaceae
Sibling taxa
A. menziesii
Subordinate taxa
A. laevisphaera, A. menziesii
Synonyms Glyphocarpa laevisphaera
Name authority (Taylor) Flowers: in A. J. Grout, Moss Fl. N. Amer. 2: 155. (1935) Schimper: Syn. Musc. Eur. ed. 2, CIII, 513. (1876)
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