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Great Basin bulrush, Nevada bulrush, Nevada clubrush

clubrush, Nevada bulrush, Nevada clubrush

Habit Herbs, perennial, internally mostly solid, without evident air cavities, cespitose or not, rhizomatous.
Rhizomes

1–4 mm diam., tough, hard.

Culms

ridged, 10–70 cm × 0.5–2 mm, hard, without evident internal air cavities.

solitary or not, ± terete, tough, wiry.

Leaves

5–10;

sheaths loose, the proximal often disintegrating to prominent fibers, often stramineous or brownish, papery, fronts membranous-hyaline with delicate veins, summit slightly concave with V-shaped veinless area and often disintegrating;

blades 0.5–1 times as long as culms, hard, without evident internal air cavities;

distal blades 3–30 cm × 0.5–2 mm, longer than sheaths, margins sparsely antrorsely spinulose or papillose, apex sharply acute.

all basal;

sheaths often disintegrating into fibers;

ligules ciliate;

blades strongly C-shaped in cross section to subcylindric, tough, wiry.

Inflorescences

proximal involucral bract 1–15 cm, resembling foliage leaf blade.

terminal, often pseudolateral, capitate;

spikelets 1–6(–10);

involucral bracts 1–3, spreading or erect, leaflike.

Spikelets

ovoid to lanceoloid or terete, 5–20 × 3–5 mm;

scales pale to dark red-brown, midribs usually stramineous, smooth;

proximal 1 or 2 scales often resembling involucral bracts, with awnlike blades, to 15 mm;

other scales in proximal part of spikelet prominently 9-veined, ovate, 4 × 3 mm, papery to often cartilaginous, margins hyaline, ciliolate, apex entire, acute to rounded.

5–20 × 3–5 mm;

scales 30–60, spirally arranged, each subtending flower, smooth, glabrous, margins ciliolate.

Flowers

perianth bristles pale brown, unequal.

bisexual;

perianth of 1–6 bristles, straight, not longer than achene, retrorsely spinulose;

stamens 3;

styles deciduous, linear, 2-fid.

Achenes

greenish to orange-brown, broadly obovoid, 2–2.3 × 1.5–1.7 mm.

plano-convex or unequally biconvex.

Amphiscirpus nevadensis

Amphiscirpus

Phenology Fruiting summer.
Habitat Sunny, saline, often alkaline, seasonally wet places
Elevation 400–2400 m (1300–7900 ft)
Distribution
from FNA
CA; CO; MT; ND; NE; NV; OR; UT; WA; WY; AB; BC; SK; South America (Argentina, Chile)
[WildflowerSearch map]
North America; South America
Discussion

Amphiscirpus nevadensis superficially resembles some dwarfed forms of Schoenoplectus pungens, with which it sometimes grows; S. pungens is readily distinguished by its trigonous culms and leaf blades, prominently 2-fid, awned scales, and beaked achenes. Amphiscirpus nevadensis differs from all North American species of Schoenoplectus in its wiry culms and leaves, prominently ciliate ligules, absence of evident internal aerenchyma, and beakless achenes. It has been reported from Delta, Manitoba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 1.

Amphiscirpus was segregated from Scirpus (A. A. Oteng-Yeboah 1974) mainly on the basis of the culm anatomy, including the absence of internal air cavities. It differs from all species of Schoenoplectus in its prominently ciliate ligules. Amphiscirpus was treated within the high Andean Phylloscirpus (J. J. Bruhl 1995) and also as a distinct, monotypic genus (P. Goetghebeur and D. A. Simpson 1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 28. FNA vol. 23, p. 27. Author: S. Galen Smith.
Parent taxa Cyperaceae > Amphiscirpus Cyperaceae
Subordinate taxa
A. nevadensis
Synonyms Scirpus nevadensis
Name authority (S. Watson) Oteng-Yeboah: Notes Roy. Bot. Gard. Edinburgh 33: 308. (1974) Oteng-Yeboah: Notes Roy. Bot. Gard. Edinburgh 33: 308. (1974)
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