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European beachgrass, European marram grass, marramgrass

American Beach grass, beach grass

Culms

(50)60-120 cm.

50-130 cm.

Panicles

12-35 cm long, 15-20 mm wide, stramineous.

9-40 cm long, 10-25 mm wide, stramineous.

Spikelets

12-14 mm.

Glumes

equaling or exceeding the florets;

callus hairs 2-6 mm;

lemmas 8-12 mm, apices shortly bifid, midveins slightly excurrent;

anthers 4-7 mm.

8-15 mm, exceeding the florets;

callus hairs 1-3 mm;

lemmas 8-11.5(14) mm, somewhat thicker than the glumes, apices acute, entire, unawned, rarely mucronate, mucros to 0.2 mm, subterminal;

anthers 3-7 mm.

Ligules

10-35 mm, acute, bifid or lacerate;

blades to 60(90) cm long and 6 mm wide when flat, usually tightly convolute and sharply pointed.

1-3 (4.6) mm, truncate to obtuse, ciliolate;

blades 15-80 cm long, 4-8 mm wide when flat, usually involute, 0.5-2.5 mm in diameter when involute;

flag leaf blades 5-39 cm.

2n

= 14, 28, 56.

Ammophila arenaria

Ammophila breviligulata

Distribution
from FNA
CA; MD; OR; PA; WA; HI; BC; NS
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CT; DE; IL; IN; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; OR; PA; RI; SC; VA; VT; WA; WI; BC; NB; NL; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Ammophila arenaria is a European species that has become naturalized in most temperate countries. It was introduced along the Pacific coast and in the interior of western North America as a sand binder. It is also reported to occur, as an introduction, on the east coast; the report has not been verified. It is known from a single 1941 collection on a sand dune in Erie County, Pennsylvania. Ammophila arenaria flowers from the end of June to August. It forms a strong association with the fungus Psilocybe azurescens, which grows, often prolifically, along the northern Oregon coast.

North American plants belong to Ammophila arenaria (L.) Link subsp. arenaria, in which the glumes exceed the lemma and the callus hairs are about 2-3 mm long. It is native from northern and western Europe to northwestern Spain. Ammophila arenaria subsp. arundinacea H. Lindb. has glumes that equal the lemma, and callus hairs 4-6 mm long. It is native around the Mediterranean.

In Europe, Ammophila arenaria hybridizes with Calamagrostis. The hybrids, which are sterile, have also been used as sand binders but not, so far as is known, in the Flora region. They differ from A. arenaria in having a less dense panicle, slender lemma awns up to 2 mm long, and callus hairs 1/2 as long as the lemmas or longer. In the Flora region, a hybrid with C. nutkaensis was reported in the 1970s from the vicinity of Newport-Waldport in coastal Oregon, where both species occur (K.L. Chambers, pers. comm.). There is no voucher to support this report.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The two subspecies overlap in their morphological characteristics and occupy similar habitats, the primary difference between them being their flowering time.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering late July to September; panicles (9.5)13-40 cm long; anthers 3.5-7 mm long
subsp. breviligulata
1. Flowering late June to early July; panicles 9-17 cm long; anthers 3-4.5 mm long
subsp. champlainensis
Source FNA vol. 24, p. 777. FNA vol. 24, p. 777.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Ammophila Poaceae > subfam. Pooideae > tribe Poeae > Ammophila
Sibling taxa
A. breviligulata
A. arenaria
Subordinate taxa
A. breviligulata subsp. breviligulata, A. breviligulata subsp. champlainensis
Name authority (L.) Link Fernald
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