Amelichloa brachychaeta |
Amelichloa |
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Puna needlegrass |
needlegrass |
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Habit | Plants with knotty, shortly rhizomatous bases. | Plants perennial; cespitose. | ||||||||
Culms | 40-90 cm tall, 1-2(3) mm thick, erect, glabrous; nodes usually 3. |
erect, with 2-3 nodes, not branching at the upper nodes; basal branching intravaginal; prophylls concealed by the leaf sheaths, winged over the keels, apices bifid, teeth 0.5-3.5 mm. |
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Leaves | mostly basal; sheaths open, smooth, glabrous; cleistogenes often present, spikelets of cleistogenes 0.5-1 mm long, with thin glumes shorter than the florets, florets unawned or with reduced awns; auricles absent; ligules scarious, rounded to acute, ciliate; blades stiff, involute, apices stiff, brown, sharply pointed, blades of the flag leaves 5-13 cm long, bases similar in width to the top of the sheaths. |
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Panicles | 10-25 cm long, 1-4 cm wide, bases sometimes included in the upper leaf sheaths; branches ascending to spreading, longest lower branches 4-12 cm. |
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Inflorescences | panicles, the main panicle terminal, apparently wholly chasmogamous. |
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Spikelets | with 1 floret; disarticulation above the glumes, beneath the floret. |
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Glumes | subequal, 6-8 mm, linear-lanceolate, 1-3-veined, midveins smooth, scabridulous, or with stiff hairs, varying within a panicle, apices acuminate; florets 4-5.5 mm long, about 0.8 mm thick, fusiform; calluses 0.4-0.5 mm, blunt, strigose, hairs 0.5-0.8 mm; lemmas pubescent over and between the veins on the proximal 1/2 at least initially, hairs 0.5-0.8 mm, sometimes glabrous at maturity between the midveins and lateral veins, distal portion glabrous, tapering to the apices, apices with 0.7-1 mm hairs around the base of the awn; awns 10-18 mm, glabrous or scabrous, usually once-geniculate; paleas 3/4 - 9/10 as long as the lemmas, pubescent over the central portion, apices involute; lodicules 3; anthers 2-2.4(3) mm, penicillate. |
exceeding the floret, acute to acuminate, 1-5-veined; florets fusiform, terete; calluses antrorsely strigose, blunt; lemmas pubescent, often more densely and/or more persistently so over the midvein and lateral veins, hairs on the proximal portion about 0.7-2 mm, hairs on the distal portion often longer; crowns not developed; awns once- or twice-geniculate, scabrous, persistent; paleas 3/4 as long as to almost equaling the lemmas, flat, hairy, hairs 0.2-1 mm, veins terminating at or near the apices, apices similar in texture to the body; lodicules 3; anthers 3, anthers sometimes all of equal size and more than 2 mm, sometimes 1 longer than 2 mm and 2 much shorter, sometimes all shorter than 2 mm; ovaries glabrous; styles with 2 branches, united at the base, stigmas plumose. |
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Caryopses | 2-3 mm long, 0.9-1 mm thick, fusiform; style bases straight, centric. |
obovoid, with 3 smooth, longitudinal ribs at maturity, stylar bases 1-2 mm, persistent, sometimes eccentric; hila linear, about as long as the caryopses. |
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Basal | sheaths mostly glabrous, margins ciliate distally; collars glabrous, often with tufts of hair to 1.5 mm on the sides; ligules 0.2-0.6 mm, membranous, strigose, ciliate, cilia to 2 mm, slightly longer at the leaf margins; blades 8-35 cm long, usually convolute and 0.5-0.8 mm in diameter, 2-3 mm wide when flat, erect, abaxial surfaces smooth, adaxial surfaces usually glabrous. |
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x | = unknown. |
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2n | = unknown. |
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Amelichloa brachychaeta |
Amelichloa |
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Distribution | ||||||||||
Discussion | Amelichloa brachychaeta has been found at a few locations in California, where it is listed as a noxious weed. It is native to Uruguay and Argentina. It is avoided by cattle because of its sharply pointed leaves. The cleistogamous panicles, which may be at or below ground level, remain a source of seeds unless the plants are completely uprooted. Amelichloa caudata and A. clandestina (see below) are a greater problem in this regard, because they appear to produce such panicles more frequently. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amelichloa includes five species, four of which are South American. The fifth species, A. clandestina, grows in northern Mexico. Two species are established in the Flora region. A third species, A. caudata. was found on ballast dumps near Portland, Oregon, at the turn of the twentieth century; it is not established in the region.Cattle avoid species of Amelichloa because of their sharply pointed leaves. This means that any of the species could become a serious problem in rangelands. Mowing favors their establishment and spread because it does not eliminate, and may disperse, the cleistogenes. The species are eaten by goats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 182. | FNA vol. 24, p. 181. | ||||||||
Parent taxa | ||||||||||
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Synonyms | Achnatherum brachychaetum | |||||||||
Name authority | (Godr.) Arriaga & Barkworth | Arriaga & Barkworth | ||||||||
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