Amelanchier utahensis |
Amelanchier interior |
|
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pale leaf serviceberry, pale serviceberry, Utah service-berry, Utah serviceberry or shadbush |
amélanchier de l'intérieur, inland serviceberry, Wiegand's shadbush |
|
Habit | Shrubs, 0.5–5 m. Stems 1–100, often colonial, much branched. | Shrubs or trees, 1–10 m. Stems 1–10, forming colonies or solitary. |
Leaves | mostly or fully unfolded; petiole (3–)6–13(–22) mm; blade suborbiculate to oval or obovate, (14–)21–36(–63) × (9–)16–32(–54) mm, base usually rounded to subcordate, sometimes cuneate, each margin with 0–3(–6) teeth on proximal 1/2 and (0–)3–5(–7) teeth in distalmost cm, largest teeth more than 1 mm, apex usually rounded to truncate or emarginate, sometimes acute and mucronate, abaxial surface moderately (sparsely or densely) hairy by flowering, sparsely to moderately hairy later, adaxial glabrous or sparsely (moderately) hairy later. |
not fully unfolded; petiole 10–30 mm; blade broadly ovate to elliptic, 30–70 × 20–50 mm, base rounded to subcordate, each margin with 3–15 teeth on proximal 1/2 and (2–)4 or 5(–7) teeth in distalmost cm, largest teeth less than 1 mm, apex short acuminate to apiculate, abaxial surface glabrous or sparsely hairy by flowering, surfaces glabrous later. |
Inflorescences | (4–)6–10(–13)-flowered, (8–)16–30(–43) mm. |
4–12-flowered, 30–75 mm. |
Pedicels | (0 or)1 or 2(or 3) subtended by a leaf, proximalmost (2–)7–16(–25) mm. |
1 or 2 subtended by a leaf, proximalmost 10–45 mm. |
Flowers | sepals usually recurved after flowering, (1.6–)2.6–4.2(–6.5) mm; petals oblanceolate to oblong, (4.9–)7.2–9.8(–14) × (1.6–)2.6–3.1(–5.3) mm; stamens (9–)13–19(–20); styles (2 or)3 or 4, (1.7–)2.3–3(–3.9) mm; ovary apex moderately to densely hairy (or glabrous). |
sepals recurved after flowering, 2–5 mm; petals obovate, 6–15 × 4–5 mm; stamens 20; styles 5, 3–5 mm; ovary apex densely (moderately) hairy (or glabrous). |
Pomes | purplish black, 6–10 mm diam. 2n = 4x. |
purple-black, 6–8 mm diam. 2n = 4x. |
Amelanchier utahensis |
Amelanchier interior |
|
Phenology | Flowering Apr–May; fruiting Jul–Sep. | Flowering May–Jun; fruiting Jul–Aug. |
Habitat | Dry rocky slopes, canyons, stream banks, mountainsides, foothills, deserts | Dry woods, bluffs rocky areas and slopes, stream banks, fields, thickets, and sandy areas, sometimes wetlands |
Elevation | 900–3500 m (3000–11500 ft) | 0–300 m (0–1000 ft) |
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; TX; UT; WA; WY; Mexico (Baja California)
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IA; IL; MI; MN; OH; SD; WI; NB; NF; NS; ON; PE; QC
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Discussion | The sparsely to moderately hairy mature leaves and twigs, rounded to truncate or emarginate leaf apices, relatively short petals, and reduced numbers of stamens and styles are distinctive characteristics for the wide-ranging and common Amelanchier utahensis. Within 25 years of its publication, about a dozen names were published that G. N. Jones (1946) considered synonyms of A. utahensis. Some recent floras have recognized some of these synonymized taxa, including A. bakeri Greene, A. covillei, A. mormonica C. K. Schneider, A. oreophila A. Nelson, and A. venulosa Greene. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amelanchier interior is distinguished by its capacity to grow into a tree to ten meters and by having sparsely hairy young leaves that are often reddish and densely hairy ovary apices. E. L. Nielsen (1939) differentiated A. interior and A. wiegandii on the basis of leaves being carinate in A. wiegandii as opposed to flat in A. interior, and leaf sinuses rounded versus acute. G. N. Jones (1946) considered those differences to be slight, and he included A. wiegandii in A. interior. The authors follow Jones, but retain the common name Wiegand’s shadbush to commemorate Wiegand’s early insights about the taxonomy of Amelanchier. M. L. Fernald (1950) considered A. wiegandii as suggesting a small-leaved A. laevis but with shorter buds, fewer teeth, fewer veins, and summit of ovary heavily tomentose. P. Landry (1975) thought A. wiegandii to be the hybrid of A. arborea and A. sanguinea, and E. G. Voss (1972–1996, vol. 2) reasoned that A. interior is a hybrid swarm involving A. laevis (or sometimes A. arborea) and plants of the A. spicata and/or A. sanguinea complex. A hybrid origin of A. interior from A. laevis and A. sanguinea is reasonable given that stem height, the number of leaf teeth, and petal length are more or less intermediate between those two species. DNA sequences from ITS region indicate that A. interior (A. wiegandii) is a possible later-generation hybrid involving a member of the western North American ITS clade (which also includes A. humilis and A. sanguinea of eastern North America) and some eastern North American taxon (C. S. Campbell et al. 1997). Multiple hybrid origins, possibly from different species, may explain the variability of A. interior. Amelanchier interior has unusually large ranges of lengths for proximal pedicels, sepals, and petals. The leaves of A. wiegandii were described by Nielsen as bronze at flowering; Jones described the leaves of A. interior as green when young. The authors assume that A. interior as they interpret it is polymorphic for the color of young leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 650. | FNA vol. 9, p. 655. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Amelanchier | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Amelanchier |
Sibling taxa | ||
Synonyms | A. covillei, A. glabra, A. gracilis, A. prunifolia, A. utahensis subsp. covillei, A. utahensis var. covillei | A. wiegandii |
Name authority | Koehne: Gatt. Pomac., 25, plate 2, fig. 20e. (1890) | E. L. Nielsen: Amer. Midl. Naturalist 22: 185, plate 13. (1939) |
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