Amelanchier utahensis
Amelanchier bartramiana
pale leaf serviceberry, pale serviceberry, Utah service-berry, Utah serviceberry or shadbush
amélanchier de Bartram, Bartram's serviceberry, mountain serviceberry, mountain shadbush, mountain shadbush or serviceberry, oblongfruit serviceberry
mostly or fully unfolded;
petiole (3–)6–13(–22) mm;
blade suborbiculate to oval or obovate, (14–)21–36(–63) × (9–)16–32(–54) mm, base usually rounded to subcordate, sometimes cuneate, each margin with 0–3(–6) teeth on proximal 1/2 and (0–)3–5(–7) teeth in distalmost cm, largest teeth more than 1 mm, apex usually rounded to truncate or emarginate, sometimes acute and mucronate, abaxial surface moderately (sparsely or densely) hairy by flowering, sparsely to moderately hairy later, adaxial glabrous or sparsely (moderately) hairy later.
half-unfolded;
petiole (2–)4.5–10.5(–25) mm;
blade narrowly elliptic-oval to oblong to broadly oval, (26–)37–51(–74) × (12–)20–29(–48) mm, base usually cuneate, each margin (2–)8–14(–27) teeth on proximal 1/2 and (2–)7–12(–21) teeth in distalmost cm, largest teeth less than 1 mm, apex acute to rounded, abaxial surface sparsely (moderately) hairy (or glabrous) by flowering, glabrous or sparsely hairy later, adaxial glabrous (or sparsely hairy) later.
(4–)6–10(–13)-flowered, (8–)16–30(–43) mm.
(1 or)2 or 3(or 4)-flowered, (6–)13–25(–38) mm.
(0 or)1 or 2(or 3) subtended by a leaf, proximalmost (2–)7–16(–25) mm.
(0 or)1(–3) subtended by a leaf, proximalmost (4–)11–21(–35) mm.
sepals usually recurved after flowering, (1.6–)2.6–4.2(–6.5) mm;
petals oblanceolate to oblong, (4.9–)7.2–9.8(–14) × (1.6–)2.6–3.1(–5.3) mm;
stamens (9–)13–19(–20);
styles (2 or)3 or 4, (1.7–)2.3–3(–3.9) mm;
ovary apex moderately to densely hairy (or glabrous).
sepals ascending to recurved after flowering, (1.7–)2.7–3.9(–6) mm;
petals oblong-oval to broadly elliptic, (5.5–)7.1–8.7(–16.9) × (2.6–)3.9–5.3(–7) mm;
stamens (8–)18–21(–25);
styles (3 or)4 or 5, (2.7–)3.8–5.2(–6.1) mm;
ovary apex densely hairy (or glabrous).
purplish black, 6–10 mm diam. 2n = 4x.
dark purple, pear-shaped, 10–15 mm diam. 2n = 2x, 3x, 4x.
Amelanchier utahensis
Amelanchier bartramiana
The sparsely to moderately hairy mature leaves and twigs, rounded to truncate or emarginate leaf apices, relatively short petals, and reduced numbers of stamens and styles are distinctive characteristics for the wide-ranging and common Amelanchier utahensis. Within 25 years of its publication, about a dozen names were published that G. N. Jones (1946) considered synonyms of A. utahensis. Some recent floras have recognized some of these synonymized taxa, including A. bakeri Greene, A. covillei, A. mormonica C. K. Schneider, A. oreophila A. Nelson, and A. venulosa Greene.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Amelanchier bartramiana is the only North American Amelanchier with leaves that are imbricate in bud, usually fewer than four flowers per inflorescence, and conic ovary apices (and hence fruits that are more or less pear-shaped rather than globose as in other members of the genus). Because of these differences, P. Landry (1975) placed A. bartramiana in its own subgenus, and all other members of the genus in another. W. H. Blanchard (1907), W. A. Robinson and C. R. Partanen (1980), and Robinson (1982) also recognized the distinctness of this species. Amelanchier bartramiana grows farther north than any other shadbush in eastern North America, and more than other shadbushes, it occupies relatively undisturbed habitats, such as peatlands and natural breaks in mature forests. Some plants in this species produce seed sexually (C. S. Campbell et al. 1987). Sexual plants of A. bartramiana are self-incompatible diploids; a tetraploid individual has been reported (A. C. Dibble et al. 1998); it had relatively large petals and might have been an autotetraploid.
Amelanchier bartramiana usually flowers with A. laevis, and it frequently hybridizes with other members of the genus. M. L. Fernald (1950) and L. Cinq-Mars (1971) reported hybrids with A. arborea (A. ×quinti-martii Louis-Marie), A. canadensis, A. fernaldii, A. gaspensis, A. humilis, A. intermedia, A. laevis, A. sanguinea, A. spicata, and A. interior. The hybrid with A. laevis can usually be found when these two species grow together (J. E. Weber and C. S. Campbell 1989). The authors have documented a hybrid between A. arborea and A. bartramiana in eastern Pennsylvania (M. B. Burgess et al., unpubl.).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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