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dwarf coastweed, pineapple weed

Habit Annuals, perennials, subshrubs, or shrubs, 1–200 cm.
Leaves

1–3+ cm, often ± fleshy.

basal, basal and cauline, or mostly cauline; mostly opposite (Lasthenia) or mostly alternate; usually sessile, sometimes obscurely petiolate;

blades (often 1–2 times pinnately lobed) or lobes often linear, ultimate margins entire or toothed, faces often ± woolly to tomentose, sometimes glabrate or glabrous, often gland-dotted.

Involucres

ovoid or obconic to campanulate or hemispheric.

Receptacles

flat, convex, hemispheric, or conic (smooth, knobby, or pitted, glabrous or hairy), usually epaleate (paleae usually 0, rare in Eriophyllum).

Ray florets

0 or 4–21, pistillate, fertile (3–8 peripheral florets pistillate, fertile, corollas tubular in Amblyopappus and Monolopia congdonii);

corollas yellow to orange, often darker proximally, sometimes purplish (usually ± bilabiate in Monolopia).

Disc florets/Disc corollas

0.6–1 mm.

2–300, bisexual, fertile;

corollas yellow to orange, tubes shorter than or about equaling funnelform or campanulate throats, lobes 4–5, deltate, glabrous or papillate;

anther thecae usually pale;

stigmatic papillae in 2 lines.

Phyllaries

persistent, mostly 3–18 in 1–2 series, (erect or reflexed in fruit) distinct or connate, mostly elliptic, lanceolate, ovate, or obovate, usually ± equal, mostly herbaceous, sometimes indurate (at least proximally), flat or weakly cupped at bases, sometimes scarious-margined, often woolly to tomentose, sometimes glabrate or glabrous.

Calyculi

0.

Heads

radiate, discoid, or disciform, borne singly or in corymbiform, glomerate, or paniculiform arrays.

Cypselae

black, 1.5–2 mm;

pappi of white or purplish, apically ± muticous scales 0.5–0.8 mm.

clavate or obovoid to terete, or obpyramidal, sometimes compressed or obcompressed, glabrous, hairy, or papillate (compressed, callous-margined, and ciliolate in Eatonella, Lasthenia chrysantha, and Monolopia congdonii; sometimes winged in Monolopia);

pappi 0 or of 1–12+ aristate, erose, laciniate, or truncate scales or awns in 1–2 series (often 2 sorts of scales in combination on 1 cypsela).

Ray

or pistillate corollas 0.6–1 mm (obscurely zygomorphic).

2n

= 16.

Amblyopappus pusillus

Asteraceae tribe Heliantheae subtribe Baeriinae

Phenology Flowering spring.
Habitat Coastal bluffs, dunes, beaches
Elevation 0–40+ m (0–100+ ft)
Distribution
from FNA
CA; South America; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
w North America; Mexico; w South America
Discussion

Genera 9, species 44 (7 genera, 41 species in the flora).

Members of Baeriinae are found mostly in western North America; there are disjuncts in western South America. H. Robinson (1981) treated Baeriinae as a relatively isolated element among epaleate subtribes of Heliantheae. B. G. Baldwin (in Baldwin et al. 2002) included Baeriinae within Madieae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves all or mostly opposite
Lasthenia
1. Leaves mostly alternate (proximal sometimes opposite)
→ 2
2. Leaves glabrous (often granular-glandular, not woolly)
Amblyopappus
2. Leaves ± woolly or tomentose (usually stems and/or phyllaries as well)
→ 3
3. Heads ± disciform (4–7 peripheral florets pistillate, corollas with laminae ca. 0.5 mm)
Monolopia
3. Heads usually radiate, sometimes discoid
→ 4
4. Phyllaries usually reflexed in fruit (ray laminae 2–2.5 mm; pappi of 2, ± lacini-ate to nearly entire, often uniaristate scales 1.5–3 mm)
Eatonella
4. Phyllaries usually ± erect in fruit
→ 5
5. Ray florets 7–11 (corollas obscurely bilabiate, laminae each opposed by adaxial lobe or tooth)
Monolopia
5. Ray florets 0 or 3–13 (corollas not bilabiate)
→ 6
6. Annuals; ray cypselae obcompressed (disc corollas with rings of hairs at bases of limbs)
Pseudobahia
6. Annuals, perennials, subshrubs, or shrubs; ray cypselae usually prismatic, 4–5-angled (disc corollas without rings of hairs)
→ 7
7. Annuals, perennials, subshrubs, or shrubs; phyllaries 4–13(–15) in 1+ series; pappi 0, or coroniform, or of 6–12+ (distinct) scales (in 1–2 similar or contrasting series)
Eriophyllum
7. Subshrubs; phyllaries 8–16+ in ± 2 series; pappi of 2–6+ (unequal or 2 opposite, longer, and ± equal, basally connate, oblong to subulate) scales (tips acute to erose)
Constancea
Source FNA vol. 21, p. 348. FNA vol. 21, p. 335.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Baeriinae > Amblyopappus Asteraceae > tribe Heliantheae
Subordinate taxa
Amblyopappus, Constancea, Eatonella, Eriophyllum, Lasthenia, Monolopia, Pseudobahia
Synonyms subtribe Eriophyllinae
Name authority Hooker & Arnott: J. Bot. (Hooker) 3: 321. (1841) Bentham & Hooker f.: Gen. Pl. 2: 200. (1873)
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