Amaranthus spinosus |
Amaranthus californicus |
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spiny amaranth, thorny amaranth |
California amaranth, California pigweed, Californian amaranth |
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Habit | Plants glabrous or sparsely pubescent in the distal younger parts of stems and branches. | Plants annual, glabrous. |
Stems | erect or sometimes ascending proximally, much-branched and bushy, rarely nearly simple, 0.3–1(–2) m; each node with paired, divergent spines (modified bracts) to 1.5(–2.5) cm. |
prostrate, whitish or tinged with red, much-branched from base, 0.1–0.5 m, rather fleshy. |
Leaves | petiole ± equaling or longer than blade; blade rhombic-ovate, ovate, or ovate-lanceolate, 3–10(–15) × 1.5–6 cm, base broadly cuneate, margins entire, plane or slightly undulate, apex acute or subobtuse to indistinctly emarginate, mucronulate. |
petiole 1/2 or less as long as blade; blade pale green, veins prominent, obovate, spatulate, or oblanceolate to linear, 0.3–2(–3) × 0.2–1.5 cm, base cuneate, margins entire, plane or slightly undulate, apex obtuse to subacute, with prominent mucro. |
Bracts | of pistillate flowers lanceolate to ovate-lanceolate, shorter than tepals, apex attenuate. |
of pistillate flowers linear, 0.5–1 mm, ± equaling tepals. |
Inflorescences | simple or compound terminal staminate spikes and axillary subglobose mostly pistillate clusters, erect or with reflexed or nodding tips, usually green to silvery green. |
axillary clusters borne from bases to tops of plants. |
Staminate flowers | often terminal or in proximal glomerules; tepals 5, equal or subequal, 1.7–2.5 mm; stamens 5. |
intermixed with pistillate; tepals (2–)3; stamens 3. |
Pistillate flowers | tepals 5, obovate-lanceolate or spatulate-lanceolate, equal or subequal, 1.2–2 mm, apex mucronate or short-aristate; styles erect or spreading; stigmas 3. |
tepals 1–3, narrowly lanceolate, unequal, usually with only 1 well-developed tepal, largest 1–1.2 mm, apex acute to acuminate; stigmas 3. |
Seeds | black, lenticular or subglobose-lenticular, 0.7–1 mm diam., smooth, shiny. |
very dark reddish brown, lenticular, (0.6–)0.7–1 mm diam., shiny. |
Utricles | ovoid to subglobose, 1.5–2.5 mm, membranaceous proximally, wrinkled and spongy or inflated distally, irregularly dehiscent or indehiscent. |
subglobose, 1–1.2 mm, smooth or wrinkled (especially in dry plants), dehiscence regularly circumscissile or tardily dehiscent. |
Amaranthus spinosus |
Amaranthus californicus |
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Phenology | Flowering summer–fall. | Flowering summer–fall. |
Habitat | Waste places, fields, roadsides, railroads, barnyards, overgrazed pastures, other disturbed habitats | Seasonally moist flats, shores of water bodies, waste places, other disturbed habitats |
Elevation | 0-700 m (0-2300 ft) | 0-2800 m (0-9200 ft) |
Distribution |
AL; AR; CA; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; MB; ON; Mexico; Central America; South America; West Indies [Introduced in North America; introduced nearly worldwide]
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CA; ID; KS; MT; NE; NV; OR; SD; TX; UT; WA; WY; AB; SK
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Discussion | Amaranthus spinosus is native to lowlands in tropical America; at present it is a pantropical weed that also occurs in some warm-temperate regions. Amaranthus spinosus, or its ancestral taxon, probably gave rise to the allopolyploid A. dubius by hybridization with some species of the A. hybridus aggregate (see above). Section Centrusa probably occupies a basal position, at least for the clade of subg. Amaranthus sect. Amaranthus, and probably for some representatives of subg. Acnida as currently outlined. Recent results of sequencing the ITS region (including ITS-1, 5.8S rDNA, and ITS-2) of nuclear ribosomal DNA from 15 species of Amaranthus occurring in China also suggest the basal position of A. spinosus among the studied species (Song B. H. et al. 2000). These results also confirm a profound divergence between subgenera Amaranthus and Albersia; the latter is called “sect. Paucestamen by the above authors. Data on the electrophoretic variation of seed proteins (R. H. Sammour et al. 1993) are also in accord with the segregation of these two subgenera; in the cited article, these groups are called sect. Amaranthus and sect. Blitopsis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4. | FNA vol. 4, p. 431. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Amaranthus | Amaranthaceae > Amaranthus > subg. Albersia |
Sibling taxa | ||
Synonyms | Mengea californica | |
Name authority | Linnaeus: Sp. Pl. 2: 991. (1753) | (Moquin-Tandon) S. Watson: in W. H. Brewer et al., Bot. California 2: 42. (1880) |
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