Amaranthus palmeri |
Amaranthus tuberculatus |
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carelessweed, Palmer's amara nth, Palmer's pigweed |
common waterhemp, rough-fruit amaranth, rough-fruit water-hemp, tall water-hemp |
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Habit | Plants glabrous or nearly so. | |
Stems | erect, branched, usually (0.3–)0.5–1.5(–3) m; proximal branches often ascending. |
erect to sometimes ascending or rarely prostrate, branched, rarely simple, usually (0.5–)1–2(–3) m. |
Leaves | long-petiolate; blade obovate or rhombic-obovate to elliptic proximally, sometimes lanceolate distally, 1.5–7 × 1–3.5 cm, base broadly to narrowly cuneate, margins entire, plane, apex subobtuse to acute, usually with terminal mucro. |
petiole 1/4–1/2 length of blade; blade ovate or obovate proximally, oblong or elliptic to narrowly lanceolate distally, 1.5–15 × 0.5–3 cm, base cuneate, margins entire, plane, apex obtuse or rounded to acute. |
Bracts | of pistillate flowers with long-excurrent midrib, 4–6 mm, longer than tepals, apex acuminate or mucronulate; of staminate flowers, 4 mm, equaling or longer than outer tepals, apex long-acuminate. |
of pistillate flowers 1–2 mm; of staminate flowers with inconspicuous to prominent midrib, 1–2 mm, apex acuminate to short-subulate. |
Inflorescences | terminal, linear spikes to panicles, usually drooping, occasionally erect, especially when young, with few axillary clusters, uninterrupted or interrupted in proximal part of plant. |
terminal, linear spikes to panicles, occasionally interrupted-moniliform, remote, globose glomerules. |
Staminate flowers | tepals 5, unequal, 2–4 mm, apex acute; inner tepals with prominent midrib excurrent as rigid spine, apex long-acuminate or mucronulate; stamens 5. |
tepals 5, inner tepals with prominent midribs excurrent as rigid spines or not, equal to unequal, 2–3 mm, apex obtuse to acute or acuminate or indistinctly mucronulate; stamens 5. |
Pistillate flowers | tepals 1.7–3.8 mm, apex acuminate, mucronulate; style branches spreading; stigmas 2(–3). |
tepals absent or 1–2, often rudimentary, 1–3 mm; style branches ± erect; stigmas 3. |
Seeds | dark reddish brown to brown, 1–1.2 mm diam., shiny. |
dark reddish brown to dark brown, 0.7–1 mm diam., shiny. |
Utricles | tan to brown, occasionally reddish brown, obovoid to subglobose, 1.5–2 mm, shorter than tepals, at maturity walls thin, almost smooth or indistinctly rugose. |
dark brown to reddish brown, not ribbed, obovoid to subglobose, 1.5–2 mm, thin, almost smooth or irregularly rugose, indehiscent, irregularly dehiscent, or dehiscence regularly circumscissile. |
Amaranthus palmeri |
Amaranthus tuberculatus |
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Phenology | Flowering summer–fall, occasionally spring–winter in southern part of its native range. | Flowering summer–fall. |
Habitat | Streambanks, disturbed habitats, especially agricultural fields, railroads, waste areas, roadsides | Wet areas, such as margins of rivers, ponds, marshes, lakes, and creeks, disturbed habitats, such as agricultural fields, roadsides, and railroads |
Elevation | 100-1000 m (300-3300 ft) | 0-1000 m (0-3300 ft) |
Distribution |
AR; AZ; CA; CO; FL; GA; IL; KS; KY; LA; MA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OH; OK; PA; SC; TN; TX; UT; VA; WI; WV; ON; Mexico [Introduced Europe, Asia, and Australia]
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AL; AR; CA; CO; CT; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NM; NV; NY; OH; PA; SC; SD; TN; TX; VT; WA; WI; WV; MB; ON; PE; QC [Introduced in North America; introduced, usually not naturalized, in Europe and other continents]
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Discussion | Originally native to the North American Southwest, from southern California to Texas and northern Mexico, Amaranthus palmeri at present is a successful invasive species, which is evident from its expansion both in eastern North America and overseas. Because of its rapid spread, the distribution data presented here are probably incomplete. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
J. D. Sauer (1967b, 1972b) separated Amaranthus rudis (A. tamariscinus) as distinct from A. tuberculatus, based primarily on absence of tepals in the pistillate flowers and indehiscent fruits. Recent work by D. B. Pratt and L. G. Clark (2001) showed that those characteristics are not constant and they recognized only one polymorphic species, A. tuberculatus. Long-term observations by K. R. Robertson strongly support the inclusion of A. rudis within A. tuberculatus. Amaranthus rudis probably was originally native to the Great Plains west of the Mississippi, from Texas to Iowa. Amaranthus tuberculatus likely had a more northern range, north of Missouri and Tennessee to the Great Lakes. The emerging evolutionary differentiation between the two related taxa was erased by agriculture and human-induced introduction and invasion. Amaranthus tuberculatus has become a major weed of agricultural fields and other disturbed habitats and is now introduced in parts of North America far outside its original range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 418. | FNA vol. 4. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus | Amaranthaceae > Amaranthus > subg. Acnida > sect. Acnida |
Sibling taxa | ||
Synonyms | Acnida tuberculata, Acnida altissima, Acnida altissima var. prostrata, Acnida altissima var. subnuda, Acnida concatenata, Acnida subnuda, Acnida tamariscina, Acnida tamariscina var. concatenata, Acnida tamariscina var. tuberculata, A. ambigens, A. rudis | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 12: 274. (1877) | (Moquin-Tandon) J. D. Sauer: Madroño 13: 18. (1955) |
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