Amaranthus palmeri |
Amaranthus spinosus |
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carelessweed, Palmer's amara nth, Palmer's pigweed |
spiny amaranth, thorny amaranth |
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Habit | Plants glabrous or nearly so. | Plants glabrous or sparsely pubescent in the distal younger parts of stems and branches. |
Stems | erect, branched, usually (0.3–)0.5–1.5(–3) m; proximal branches often ascending. |
erect or sometimes ascending proximally, much-branched and bushy, rarely nearly simple, 0.3–1(–2) m; each node with paired, divergent spines (modified bracts) to 1.5(–2.5) cm. |
Leaves | long-petiolate; blade obovate or rhombic-obovate to elliptic proximally, sometimes lanceolate distally, 1.5–7 × 1–3.5 cm, base broadly to narrowly cuneate, margins entire, plane, apex subobtuse to acute, usually with terminal mucro. |
petiole ± equaling or longer than blade; blade rhombic-ovate, ovate, or ovate-lanceolate, 3–10(–15) × 1.5–6 cm, base broadly cuneate, margins entire, plane or slightly undulate, apex acute or subobtuse to indistinctly emarginate, mucronulate. |
Bracts | of pistillate flowers with long-excurrent midrib, 4–6 mm, longer than tepals, apex acuminate or mucronulate; of staminate flowers, 4 mm, equaling or longer than outer tepals, apex long-acuminate. |
of pistillate flowers lanceolate to ovate-lanceolate, shorter than tepals, apex attenuate. |
Inflorescences | terminal, linear spikes to panicles, usually drooping, occasionally erect, especially when young, with few axillary clusters, uninterrupted or interrupted in proximal part of plant. |
simple or compound terminal staminate spikes and axillary subglobose mostly pistillate clusters, erect or with reflexed or nodding tips, usually green to silvery green. |
Staminate flowers | tepals 5, unequal, 2–4 mm, apex acute; inner tepals with prominent midrib excurrent as rigid spine, apex long-acuminate or mucronulate; stamens 5. |
often terminal or in proximal glomerules; tepals 5, equal or subequal, 1.7–2.5 mm; stamens 5. |
Pistillate flowers | tepals 1.7–3.8 mm, apex acuminate, mucronulate; style branches spreading; stigmas 2(–3). |
tepals 5, obovate-lanceolate or spatulate-lanceolate, equal or subequal, 1.2–2 mm, apex mucronate or short-aristate; styles erect or spreading; stigmas 3. |
Seeds | dark reddish brown to brown, 1–1.2 mm diam., shiny. |
black, lenticular or subglobose-lenticular, 0.7–1 mm diam., smooth, shiny. |
Utricles | tan to brown, occasionally reddish brown, obovoid to subglobose, 1.5–2 mm, shorter than tepals, at maturity walls thin, almost smooth or indistinctly rugose. |
ovoid to subglobose, 1.5–2.5 mm, membranaceous proximally, wrinkled and spongy or inflated distally, irregularly dehiscent or indehiscent. |
Amaranthus palmeri |
Amaranthus spinosus |
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Phenology | Flowering summer–fall, occasionally spring–winter in southern part of its native range. | Flowering summer–fall. |
Habitat | Streambanks, disturbed habitats, especially agricultural fields, railroads, waste areas, roadsides | Waste places, fields, roadsides, railroads, barnyards, overgrazed pastures, other disturbed habitats |
Elevation | 100-1000 m (300-3300 ft) | 0-700 m (0-2300 ft) |
Distribution |
AR; AZ; CA; CO; FL; GA; IL; KS; KY; LA; MA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OH; OK; PA; SC; TN; TX; UT; VA; WI; WV; ON; Mexico [Introduced Europe, Asia, and Australia]
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AL; AR; CA; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; MB; ON; Mexico; Central America; South America; West Indies [Introduced in North America; introduced nearly worldwide]
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Discussion | Originally native to the North American Southwest, from southern California to Texas and northern Mexico, Amaranthus palmeri at present is a successful invasive species, which is evident from its expansion both in eastern North America and overseas. Because of its rapid spread, the distribution data presented here are probably incomplete. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amaranthus spinosus is native to lowlands in tropical America; at present it is a pantropical weed that also occurs in some warm-temperate regions. Amaranthus spinosus, or its ancestral taxon, probably gave rise to the allopolyploid A. dubius by hybridization with some species of the A. hybridus aggregate (see above). Section Centrusa probably occupies a basal position, at least for the clade of subg. Amaranthus sect. Amaranthus, and probably for some representatives of subg. Acnida as currently outlined. Recent results of sequencing the ITS region (including ITS-1, 5.8S rDNA, and ITS-2) of nuclear ribosomal DNA from 15 species of Amaranthus occurring in China also suggest the basal position of A. spinosus among the studied species (Song B. H. et al. 2000). These results also confirm a profound divergence between subgenera Amaranthus and Albersia; the latter is called “sect. Paucestamen by the above authors. Data on the electrophoretic variation of seed proteins (R. H. Sammour et al. 1993) are also in accord with the segregation of these two subgenera; in the cited article, these groups are called sect. Amaranthus and sect. Blitopsis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 418. | FNA vol. 4. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus | Amaranthaceae > Amaranthus > subg. Amaranthus |
Sibling taxa | ||
Name authority | S. Watson: Proc. Amer. Acad. Arts 12: 274. (1877) | Linnaeus: Sp. Pl. 2: 991. (1753) |
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