Amaranthus palmeri |
Amaranthus dubius |
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carelessweed, Palmer's amara nth, Palmer's pigweed |
spleen amaranth |
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Habit | Plants glabrous or nearly so. | Plants glabrous or sparsely pubescent in distal parts. |
Stems | erect, branched, usually (0.3–)0.5–1.5(–3) m; proximal branches often ascending. |
erect, green, branched, 0.3–1 m. |
Leaves | long-petiolate; blade obovate or rhombic-obovate to elliptic proximally, sometimes lanceolate distally, 1.5–7 × 1–3.5 cm, base broadly to narrowly cuneate, margins entire, plane, apex subobtuse to acute, usually with terminal mucro. |
petiole of proximal leaves equaling or longer than blade, becoming shorter distally; blade rhombic-ovate or ovate to elliptic, 3–12 × 2–8 cm, base broadly cuneate, margins entire, apex slightly acuminate to obtuse and faintly emarginate, mucronate. |
Bracts | of pistillate flowers with long-excurrent midrib, 4–6 mm, longer than tepals, apex acuminate or mucronulate; of staminate flowers, 4 mm, equaling or longer than outer tepals, apex long-acuminate. |
lanceolate, shorter than 2 mm, shorter than tepals, apex spinescent. |
Inflorescences | terminal, linear spikes to panicles, usually drooping, occasionally erect, especially when young, with few axillary clusters, uninterrupted or interrupted in proximal part of plant. |
terminal panicles and axillary spikes; panicles erect or often drooping, green, dense, branched, leafless at least distally. |
Staminate flowers | tepals 5, unequal, 2–4 mm, apex acute; inner tepals with prominent midrib excurrent as rigid spine, apex long-acuminate or mucronulate; stamens 5. |
usually clustered at tips of inflorescence branches, sometimes gathered in proximal glomerules (as in A. spinosus); tepals 5, equal or subequal; stamens 5. |
Pistillate flowers | tepals 1.7–3.8 mm, apex acuminate, mucronulate; style branches spreading; stigmas 2(–3). |
tepals 5, oblong-spatulate to oblong, not clawed, 1.5–2 mm, apex acute, often very shortly mucronate; style branches strongly spreading, shorter than body of fruit; stigmas 3. |
Seeds | dark reddish brown to brown, 1–1.2 mm diam., shiny. |
dark reddish brown to black, subglobose or lenticular, 0.8–1 mm diam., shiny, smooth. |
Utricles | tan to brown, occasionally reddish brown, obovoid to subglobose, 1.5–2 mm, shorter than tepals, at maturity walls thin, almost smooth or indistinctly rugose. |
ovoid or subglobose, 1.5–2 mm, slightly shorter than tepals, smooth to irregularly wrinkled, dehiscence regularly circumscissile. |
2n | = 64. |
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Amaranthus palmeri |
Amaranthus dubius |
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Phenology | Flowering summer–fall, occasionally spring–winter in southern part of its native range. | Flowering summer–fall in tropics, various seasons in subtropics. |
Habitat | Streambanks, disturbed habitats, especially agricultural fields, railroads, waste areas, roadsides | Waste places, disturbed habitats |
Elevation | 100-1000 m (300-3300 ft) | 0-100 m (0-300 ft) |
Distribution |
AR; AZ; CA; CO; FL; GA; IL; KS; KY; LA; MA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OH; OK; PA; SC; TN; TX; UT; VA; WI; WV; ON; Mexico [Introduced Europe, Asia, and Australia]
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FL; South America; West Indies [Introduced in North America; introduced and locally naturalized Europe, Asia, Africa] |
Discussion | Originally native to the North American Southwest, from southern California to Texas and northern Mexico, Amaranthus palmeri at present is a successful invasive species, which is evident from its expansion both in eastern North America and overseas. Because of its rapid spread, the distribution data presented here are probably incomplete. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amaranthus dubius, a morphologically deviant allopolyploid, is very close genetically to both A. spinosus (sect. Centrusa) and members of sect. Amaranthus. This species most probably originated as a result of ancient hybridization between A. spinosus and either A. hybridus or A. quitensis (W. F. Grant 1959; T. N. Khoshoo and M. Pal 1972; M. Pal and T. N. Khoshoo 1965; J. D. Sauer 1967b; V. Srivastava et al. 1977). Amaranthus nothosect. Dubia Mosyakin & K. R. Robertson (A. sect. Amaranthus × A. sect. Centrusa), was proposed to accommodate A. dubius (S. L. Mosyakin and K. R. Robertson 1996). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 418. | FNA vol. 4, p. 425. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus | Amaranthaceae > Amaranthus > subg. Amaranthus |
Sibling taxa | ||
Name authority | S. Watson: Proc. Amer. Acad. Arts 12: 274. (1877) | Martius ex Thellung: Fl. Adv. Montpellier 38: 203. (1912) |
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