Amaranthus palmeri |
Amaranthaceae |
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carelessweed, Palmer's amara nth, Palmer's pigweed |
amaranth family |
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Habit | Plants glabrous or nearly so. | Herbs, rarely subshrubs, annual or perennial; trichomes simple (branched in Tidestromia). | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, branched, usually (0.3–)0.5–1.5(–3) m; proximal branches often ascending. |
without nodal spines (Amaranthus spinosus sometimes with paired nodal spines). |
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Leaves | long-petiolate; blade obovate or rhombic-obovate to elliptic proximally, sometimes lanceolate distally, 1.5–7 × 1–3.5 cm, base broadly to narrowly cuneate, margins entire, plane, apex subobtuse to acute, usually with terminal mucro. |
alternate or opposite, exstipulate, usually petiolate; blade margins entire (entire or serrulate in Iresine; entire, crispate, or erose in Amaranthus). |
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Bracts | of pistillate flowers with long-excurrent midrib, 4–6 mm, longer than tepals, apex acuminate or mucronulate; of staminate flowers, 4 mm, equaling or longer than outer tepals, apex long-acuminate. |
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Inflorescences | terminal, linear spikes to panicles, usually drooping, occasionally erect, especially when young, with few axillary clusters, uninterrupted or interrupted in proximal part of plant. |
cymules arranged in spikes, panicles, thyrses, heads, glomerules, clusters, or racemes; each flower subtended by 1 bract and 2 bracteoles (latter sometimes 1 or absent in Amaranthus). |
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Flowers | bisexual or unisexual (plants then monoecious or dioecious), hypogynous, generally small or minute; tepals mostly (1–)4–5 or absent, distinct or connate into cups or tubes, scarious, chartaceous, membranaceous, or indurate; stamens 2–5, filaments basally connate into cups or tubes, rarely distinct, alternating with pseudostaminodes (appendages on staminal tubes) or not, anthers 2-locular with 1 line of dehiscence or 4-locular with 2 lines of dehiscence; ovary superior, 1-locular; ovules 1 or, rarely, 2–many; style 1 or absent; stigmas 1–3(–5). |
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Staminate flowers | tepals 5, unequal, 2–4 mm, apex acute; inner tepals with prominent midrib excurrent as rigid spine, apex long-acuminate or mucronulate; stamens 5. |
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Pistillate flowers | tepals 1.7–3.8 mm, apex acuminate, mucronulate; style branches spreading; stigmas 2(–3). |
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Fruits | utricles, dry, dehiscent or not. |
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Seeds | dark reddish brown to brown, 1–1.2 mm diam., shiny. |
black, reddish brown, or brown, lenticular, subglobose or globose (rarely cylindric), usually small; embryo peripheral, surrounding mealy perisperm. |
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Utricles | tan to brown, occasionally reddish brown, obovoid to subglobose, 1.5–2 mm, shorter than tepals, at maturity walls thin, almost smooth or indistinctly rugose. |
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Amaranthus palmeri |
Amaranthaceae |
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Phenology | Flowering summer–fall, occasionally spring–winter in southern part of its native range. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Streambanks, disturbed habitats, especially agricultural fields, railroads, waste areas, roadsides | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 100-1000 m (300-3300 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AR; AZ; CA; CO; FL; GA; IL; KS; KY; LA; MA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OH; OK; PA; SC; TN; TX; UT; VA; WI; WV; ON; Mexico [Introduced Europe, Asia, and Australia]
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Nearly worldwide; most abundant in tropics; subtropics; and warm-temperate regions; evidently absent from alpine and arctic regions |
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Discussion | Originally native to the North American Southwest, from southern California to Texas and northern Mexico, Amaranthus palmeri at present is a successful invasive species, which is evident from its expansion both in eastern North America and overseas. Because of its rapid spread, the distribution data presented here are probably incomplete. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 65, species ca. 900 (12 genera, 80 species in the flora). Centers of diversity for Amaranthaceae are southwestern North America, Central America, South America, and Africa south of the Sahara Desert. Generic limits are not well defined in some groups; fewer than 60 or more than 70 genera could be recognized. Some species occur in severe habitats such as sandy, calcareous, gypseous, saline, or serpentine soils in deserts, semideserts, and seashores. Some species are weedy, including the major agricultural weeds in Amaranthus. Some species are cultivated as ornamentals, particularly Amaranthus caudatus (love-lies-bleeding), A. hypochondriacus (prince’s-feather), A. tricolor (Joseph’s-coat), Celosia cristata (cockscomb), and Gomphrena globosa (globe-amaranth). Native Americans domesticated white-seeded grain amaranths (A. caudatus, A. cruentus, and A. hypochondriacus) for use as cereal grains. Some species of Amaranthus and Celosia are potherbs. Amaranthaceae are usually divided into subfamilies Amaranthoideae (anthers 4-locular with two lines of dehiscence) and Gomphrenoideae Schinz (anthers 2-locular with one line of dehiscence). Amaranthaceae and Chenopodiaceae have long been recognized as allied families that share a number of features: generally small flowers, one perianth whorl, a syncarpous gynoecium with a superior ovary and often only one ovule, basal or free-central placentation, pollen characteristics, centrospermous embryo development, betalain pigments, and P-type form (c) sieve-element plastids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 418. | FNA vol. 4, p. 405. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 12: 274. (1877) | Jussieu | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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