Amaranthus hybridus |
Amaranthus watsonii |
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green amaranth, green pigweed, hybrid amaranth, slender pigweed, slim amaranth, smooth amaranth, smooth pigweed |
Watson's amaranth |
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Habit | Plants glabrous or glabrescent, or distal parts of stem and branches slightly pubescent when young. | Plants pubescent and glandular-pubescent, especially on bracts. |
Stems | erect, green or sometimes reddish purple, rarely under-developed plants ascending, branched to nearly simple, 0.3–2(–2.5) m. |
ascending to erect, usually much-branched, 0.1–1 m; branches usually ascending. |
Leaves | petiole 1/2 as long as to equaling blade; blade ovate, rhombic-ovate, or lanceolate, (2–)4–15 × (1–)2–6 cm, base cuneate to broadly cuneate, margins entire, apex acute to obtuse, with mucro. |
petiole shorter than or equaling blade; blade ovate, obovate to elliptic, or oblong-elliptic, 1–8 × 0.5–4 cm, base broadly cuneate to nearly rounded, margins entire, plane or slightly undulate, apex obtuse to emarginate, usually with terminal mucro. |
Bracts | lanceolate-linear to subulate, 2–3.5(–4) mm, subequal to or 2 times as long as tepals, apex spinescent. |
of pistillate flowers with long-excurrent midrib, 3–4 mm, longer than tepals, apex long-acuminate or mucronulate; of staminate flowers 2.5–4 mm, usually equaling outer tepals, apex long-acuminate or mucronulate. |
Inflorescences | terminal and axillary, erect or reflexed, occasionally nodding, green or olive green, occasionally with silvery or reddish purple tint, leafless at least distally, terrminal inflorescence often slightly nodding with numerous shorter branches at base. |
mostly terminal, erect spikes to panicles, usually thick and uninterrupted, with few axillary clusters in basal part of plant. |
Staminate flowers | at tips of inflorescences; tepals 5; stamens (4–)5. |
tepals 5, equal or subequal, 1.5–2(–3) mm, apex acute or almost obtuse; inner tepal apex acuminate or mucronulate; stamens 3–5. |
Pistillate flowers | tepals 5, lanceolate to lanceolate-linear, subequal or unequal, 1.5–3 mm, membranaceous, apex acute or acuminate, gradually narrowing into aristate tip; style branches erect, shorter than body of fruit; stigmas 3. |
tepals with dark midribs not excurrent, spatulate to fan-shaped, 1.7–2.2 mm, apex obtuse, with terminal mucro; style branches spreading; stigmas 2(–3). |
Seeds | black to dark reddish brown, lenticular to lenticular-globose, 1–1.3 mm, smooth, shiny. |
dark reddish brown to nearly black, (0.8–)1–1.2 mm diam., shiny. |
Utricles | obovoid or elongate-ovoid, 1.5–2.5 mm, shorter than tepals, smooth proximally, lid verrucose or rugose, dehiscence regularly circumscissile, or rarely in some presumably hybrid forms, irregularly dehiscent or indehiscent. |
light brown to brown, obovoid to subglobose, 1.5–2 mm, shorter than tepals, walls thin, smooth or indistinctly rugose. |
Amaranthus hybridus |
Amaranthus watsonii |
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Phenology | Flowering summer–fall. | Flowering late winter–spring, summer–fall. |
Habitat | Waste places, agricultural and fallow fields, railroads, roadsides, riverbanks, other disturbed habitats | Coastal dunes, beaches, sandy inland areas, weakly saline flats |
Elevation | 0-2500 m (0-8200 ft) | 0-100 m (0-300 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; BC; MB; NS; ON; QC; Mexico; Central America; South America; West Indies [Widely introduced or naturalized in tropical, subtropical, and warm-temperate regions worldwide]
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AZ; CA; Mexico (Baja California, Baja California Sur, Sonora); Europe (reported as rare, non-naturalized casual alien)
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Discussion | Originally a riverside pioneer in eastern North America, now Amaranthus hybridus is extremely abundant in agricultural fields and other disturbed habitats. Related cultivated species have been reported from the flora area, including A. caudatus, A. hypochondriacus, and A. cruentus; there is no evidence that they are established; specimens identified as these species are often variants of A. hybridus. Distribution of Amaranthus hybridus in North America needs clarification because the name was misapplied to other species, notably A. powellii, and specimens of A. retroflexus, A. powellii, and A. hybridus are frequently interchangeably misidentified. Forms of A. hybridus and A. powellii with reddish inflorescences are often misidentified as escaped and hence presumably naturalized, cultivated species A. caudatus Linnaeus, A. hypochondriacus Linnaeus, and A. cruentus Linnaeus. Amaranthus hybridus is extremely variable. In particular, there are numerous North American specimens with subobtuse tepals and thick inflorescences, suggesting hybridization with A. retroflexus. In Europe such presumably hybrid forms are known as A. ×ozanonii Thellung (A. Thellung 1914–1919). A new, presumably hybridogenous taxon, Amaranthus ×tucsonensis Henrickson, was recently described from Arizona (J. Henrickson 1999). It was suggested that one of its parents is A. hybridus; the other parental species (probably a species with obtuse or spatulate tepals) remains unknown. The problem of proper taxonomic position and origin of A. ×tucsonensis needs further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4. | FNA vol. 4, p. 418. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Amaranthus | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 2: 990. (1753) | Standley: Bull. Torrey Bot. Club 41: 505. (1914) |
Web links |
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