Amaranthus hybridus |
Amaranthus arenicola |
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green amaranth, green pigweed, hybrid amaranth, slender pigweed, slim amaranth, smooth amaranth, smooth pigweed |
sand amaranth, sandhill amaranth, Sandhills pigweed |
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Habit | Plants glabrous or glabrescent, or distal parts of stem and branches slightly pubescent when young. | Plants glabrous or nearly so. |
Stems | erect, green or sometimes reddish purple, rarely under-developed plants ascending, branched to nearly simple, 0.3–2(–2.5) m. |
erect, usually branched or occasionally ± simple, 0.4–1.5(–2) m; branches sometimes ascending. |
Leaves | petiole 1/2 as long as to equaling blade; blade ovate, rhombic-ovate, or lanceolate, (2–)4–15 × (1–)2–6 cm, base cuneate to broadly cuneate, margins entire, apex acute to obtuse, with mucro. |
petiole shorter than or rarely ± equaling blade; blade mostly narrowly ovate, obovate, elliptic, or lanceolate, 1.5–8 × 0.5–3 cm, thin and soft, base cuneate to nearly rounded, margins entire, plane or irregularly undulate, apex subacute to obtuse, with terminal mucro. |
Bracts | lanceolate-linear to subulate, 2–3.5(–4) mm, subequal to or 2 times as long as tepals, apex spinescent. |
of pistillate flowers with short, excurrent midrib, (1.5–)2–2.5 mm, equaling tepals or nearly so, apex acute or acuminate; of staminate flowers with prominent midribs, 2–3.5 mm, shorter than tepals, apex acute. |
Inflorescences | terminal and axillary, erect or reflexed, occasionally nodding, green or olive green, occasionally with silvery or reddish purple tint, leafless at least distally, terrminal inflorescence often slightly nodding with numerous shorter branches at base. |
mostly terminal, spikes to panicles, erect to nodding, rarely with axillary clusters in proximal part of plant. |
Staminate flowers | at tips of inflorescences; tepals 5; stamens (4–)5. |
tepals 5, equal or subequal, 3 mm, apex obtuse to subacute; inner tepals with apex indistinctly mucronulate; stamens 5. |
Pistillate flowers | tepals 5, lanceolate to lanceolate-linear, subequal or unequal, 1.5–3 mm, membranaceous, apex acute or acuminate, gradually narrowing into aristate tip; style branches erect, shorter than body of fruit; stigmas 3. |
tepals spatulate, 1.5–2.5 mm, apex obtuse, with terminal mucro; style branches ± erect; stigmas 2–3. |
Seeds | black to dark reddish brown, lenticular to lenticular-globose, 1–1.3 mm, smooth, shiny. |
dark reddish brown, (0.9–)1–1.2 mm diam., shiny. |
Utricles | obovoid or elongate-ovoid, 1.5–2.5 mm, shorter than tepals, smooth proximally, lid verrucose or rugose, dehiscence regularly circumscissile, or rarely in some presumably hybrid forms, irregularly dehiscent or indehiscent. |
light brown to brown, subglobose, 1.5–2 mm, shorter than tepals, walls thin, usually smooth. |
Amaranthus hybridus |
Amaranthus arenicola |
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Phenology | Flowering summer–fall. | Flowering summer–fall. |
Habitat | Waste places, agricultural and fallow fields, railroads, roadsides, riverbanks, other disturbed habitats | Sandy habitats, sand hills, riverbanks, creeks, lakes, disturbed areas, agricultural fields |
Elevation | 0-2500 m (0-8200 ft) | 0-2000 m (0-6600 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; BC; MB; NS; ON; QC; Mexico; Central America; South America; West Indies [Widely introduced or naturalized in tropical, subtropical, and warm-temperate regions worldwide]
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AR; AZ; CA; CO; IA; IL; IN; KS; KY; LA; MI; MO; MT; NE; NJ; NM; NV; OK; PA; SD; TN; TX; VA; WI; WY
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Discussion | Originally a riverside pioneer in eastern North America, now Amaranthus hybridus is extremely abundant in agricultural fields and other disturbed habitats. Related cultivated species have been reported from the flora area, including A. caudatus, A. hypochondriacus, and A. cruentus; there is no evidence that they are established; specimens identified as these species are often variants of A. hybridus. Distribution of Amaranthus hybridus in North America needs clarification because the name was misapplied to other species, notably A. powellii, and specimens of A. retroflexus, A. powellii, and A. hybridus are frequently interchangeably misidentified. Forms of A. hybridus and A. powellii with reddish inflorescences are often misidentified as escaped and hence presumably naturalized, cultivated species A. caudatus Linnaeus, A. hypochondriacus Linnaeus, and A. cruentus Linnaeus. Amaranthus hybridus is extremely variable. In particular, there are numerous North American specimens with subobtuse tepals and thick inflorescences, suggesting hybridization with A. retroflexus. In Europe such presumably hybrid forms are known as A. ×ozanonii Thellung (A. Thellung 1914–1919). A new, presumably hybridogenous taxon, Amaranthus ×tucsonensis Henrickson, was recently described from Arizona (J. Henrickson 1999). It was suggested that one of its parents is A. hybridus; the other parental species (probably a species with obtuse or spatulate tepals) remains unknown. The problem of proper taxonomic position and origin of A. ×tucsonensis needs further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amaranthus arenicola is native to the central and southwestern Great Plains, from Texas to Nebraska or South Dakota, and occurs as occasionally introduced in other regions of North America and in Europe, but it is not naturalized. However, many staminate specimens of A. tuberculatus have been misidentified as this species, and some of the state references listed above may be in error. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4. | FNA vol. 4, p. 419. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Amaranthus | Amaranthaceae > Amaranthus > subg. Acnida > sect. Saueranthus |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 2: 990. (1753) | I. M. Johnston: J. Arnold Arbor. 29: 193. (1948) |
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