Amaranthus deflexus |
Amaranthus graecizans |
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Argentina amaranth, deflexed amaranth, large-fruit amaranth, low amaranth |
Mediterranean amaranth, spreading pigweed, tumbleweed |
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Habit | Plants short-lived perennial or annual, pubescent in distal parts of plant or becoming glabrescent at maturity. | Plants annual, pubescent in distal parts or becoming glabrescent at maturity. |
Stems | ascending or prostrate, profusely branched basally, radiating from rootstock, mostly 0.2–0.5 m. |
erect to ascending or decumbent, branched at or distal to base, 0.1–0.9 m. |
Leaves | petiole 1/2 as long as to equaling blade; blade rhombic-ovate or ovate to lanceolate, 1–2 × 0.5–1 cm, base tapering or cuneate, margins entire, plane or slightly undulate, apex subacute, obtuse, or retuse or shallowly emarginate, mucronulate. |
petiole variable in length; blade lanceolate to nearly linear or rhombic-ovate to elliptic-ovate, (1.5–)2–4(–5) × 1–3 cm, base cuneate to broadly cuneate, margins entire, plane, rarely indistinctly undulate, apex subacute to obtuse or emarginate, mucronulate. |
Bracts | of pistillate flowers linear, 0.5–1 mm, 1/2 as long as tepals. |
lanceolate, subspinescent, 1.5–2 mm, shorter or slightly longer than tepals. |
Inflorescences | terminal, erect, compact, pyramidal panicles and also some axillary clusters, green or silvery green, occasionally tinged with red, leafless at least distally. |
axillary glomerules, green. |
Staminate flowers | clustered at tips of inflorescences; tepals 2–3; stamens 2–3. |
intermixed with pistillate; tepals 3, equal or subequal; stamens 3. |
Pistillate flowers | tepals 2–3, narrowly elliptic or oblanceolate, not clawed, equal or subequal, 1.2–2 mm, apex broadly acute; style branches erect; stigmas 3. |
tepals 3, erect, elliptic to lanceolate-elliptic, equal or subequal, 1.5–2 mm, apex short-acuminate; style branches slightly spreading; stigmas (2–)3. |
Seeds | very dark brown to black, 1–1.2 mm diam., shiny, filling only proximal portion of fruit. |
black, lenticular, 1–1.3(–1.6) mm diam., smooth or indistinctly punctate. |
Utricles | marked with 2(–3) green lines that intersect at apex and divide fruit into halves or quarters, slightly to distinctly inflated, ellipsoid, 2–3 mm, distinctly longer than tepals, smooth (in dry plants wrinkled or rugose), indehiscent. |
subglobose to broadly elliptic, 2–2.5 mm, slightly rugose, dehiscence regularly circumscissile, rarely irregularly dehiscent. |
Amaranthus deflexus |
Amaranthus graecizans |
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Phenology | Flowering summer–fall. | Flowering summer–fall. |
Habitat | Weedy areas, ballast heaps, railroads, other disturbed habitats | On ballast |
Elevation | 0-500 m [0-1600 ft] | 0 m [0 ft] |
Distribution |
AL; CA; FL; GA; LA; MA; NJ; NY; OR; PA; TN; VA; native to South America [Introduced in North America; locally introduced or naturalized in tropical to warm-temperate regions of the globe]
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NJ; native to Eurasia (Mediterranean area, s Asia); n Africa [Introduced in North America; locally introduced in Australia]
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Discussion | The hybrid between Amaranthus deflexus and A. muricatus was described from Europe as A. ×tarraconensis Sennen & Pau (see J. L. Carretero 1979) and may be expected in North America in the future in places of possible co-occurrence of the parental species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In North America the name Amaranthus graecizans has been constantly misapplied to the common North American taxa A. albus and A. blitoides. Consequently, A. graecizans has been excluded from lists of North American plants. Recently, herbarium specimens (casual aliens collected in 1879 on ballast in Camden, New Jersey) of A. graecizans subsp. sylvestris were discovered (M. Costea et al. 2001b). Probably, the species disappeared in North America long ago, but, considering the long history of misidentification and confusion, there is also some chance that it may occur locally as an introduced species. Three subspecies are usually recognized within Amaranthus graecizans in the Old World: subsp. graecizans, subsp. sylvestris (Villars) Brenan, and subsp. thellungianus (Nevski) Gusev. Only subsp. sylvestris, characterized by rhombic-ovate to elliptic-ovate leaves (as compared to lanceolate to almost linear leaves in subsp. graecizans) and comparatively large seeds has so far been reported from North America. Despite its superficial similarity to Amaranthus albus and A. blitoides, A. graecizans seems to be more closely related to other Old World taxa with trimerous flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 430. | FNA vol. 4. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Linnaeus: Mant. Pl. 2: 295. (1771) | Linnaeus: Sp. Pl. 2: 990. (1753) |
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