Amaranthus australis |
Amaranthus retroflexus |
|
---|---|---|
southern amaranth, southern water-hemp |
common amaranth, pigweed amaranth, red-root amaranth, red-root pigweed, rough pigweed, wild-beet amaranth |
|
Habit | Plants densely to moderately pubescent, especially distal parts of stem and branches. | |
Stems | erect, branched, stout to robust, usually 1.5–3 m (occasionally to 9 m!) × 30 cm. |
erect, reddish near base, branched in distal part to simple 0.2–1.5(–2) m; underdeveloped or damaged plants rarely ascending to nearly prostrate. |
Leaves | petiole 1/3–2/3 length of blade; blade narrowly lanceolate to narrowly ovate, 10–20 × 1–4 cm, base cuneate, margins entire, plane, apex acute or long-attenuate to acuminate. |
petiole 1/2 to equaling blade; blade ovate to rhombic-ovate, 2–15 × 1–7 cm, base cuneate to rounded-cuneate, margins entire, plane or slightly undulate, apex acute, obtuse, or slightly emarginate, with terminal mucro. |
Bracts | of pistillate flowers 1.5–2 mm; of staminate flowers with moderately heavy midribs, 1.5–2 mm. |
lanceolate to subulate, (2.5–)3.5–5(–6) mm, exceeding tepals, apex acuminate with excurrent midrib. |
Inflorescences | mostly terminal, linear spikes to panicles, usually interrupted. |
terminal and axillary, erect or reflexed at tip, green or silvery green, often with reddish or yellowish tint, branched, leafless at least distally, usually short and thick. |
Staminate flowers | tepals 5, inner tepals with moderately prominent, excurrent midribs, equal, 2–2.5 mm, apex subacute to mucronulate; stamens 5. |
few at tips of inflorescences; tepals 5; stamens (3–)4–5. |
Pistillate flowers | tepals absent; style branches spreading; stigmas 3–5. |
tepals 5, spatulate-obovate, lanceolate-spatulate, not clawed, subequal or unequal, (2–)2.5–3.5(–4) mm, membranaceous, apex emarginate or obtuse, with mucro; style branches erect or slightly spreading,; stigmas 3. |
Seeds | reddish brown to dark brown, 1–1.2 mm diam., shiny. |
black to dark reddish brown, lenticular to subglobose-lenticular, 1–1.3 mm, smooth, shiny. |
Utricles | stramineous to brown, with 3(–5) longitudinal ridges corresponding to 3–5 style branches, elliptic or obovoid, 1.5–2.5 mm, slightly fleshy, smooth (slightly rugose in herbarium specimens). |
broadly obovoid to broadly elliptic, 1.5–2.5 mm, shorter than or subequal to tepals, smooth or slightly rugose, especially near base and in distal part, dehiscence regularly circumscissile. |
Amaranthus australis |
Amaranthus retroflexus |
|
Phenology | Flowering summer–fall. | Flowering summer–fall. |
Habitat | Freshwater and brackish wetland habitats, coastal marshes, swamps, riverbanks, bayous, canals, ditches, estuaries, lakeshores, hammocks | Banks of rivers, lakes, and streams, disturbed habitats, agricultural fields, railroads, roadsides, waste areas |
Elevation | 0-100 m (0-300 ft) | 0-2500 m (0-8200 ft) |
Distribution |
AL; AR; FL; GA; LA; MS; NC; TN; TX; VA; e Mexico; West Indies; n South America
|
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; SPM [Introduced and naturalized nearly worldwide]
|
Discussion | Plants of Amaranthus australis, a herbaceous annual, can be amazingly tall, with a single hollow main stem, up to 9 m, and the stem base can reach 30 cm in diameter. Large plants may somewhat resemble young trees of Taxodium distichum var. imbricarium, pondcypress. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amaranthus retroflexus, native to central and eastern North America, is a successful invasive species and has effectively colonized a wide range of habitats on all inhabited continents. Its variability is extremely wide; usually the species is easily recognized and its identification causes no specific problems. Infraspecific entities described within A. retroflexus are mostly ecologic variants of little or no taxonomic value. Two varieties are more easily recognized: the common var. retroflexus, with bracts about 1.5–2 times as long as tepals, and a more rare var. delilei (Richter & Loret) Thellung (= A. delilei Richter & Loret), with bracts 1–1.5 times as long as tepals. Occasional forms morphologically intermediate between Amaranthus retroflexus and taxa of the A. hybridus aggregate (e.g., A. powellii and A. hybridus, in the strict sense) are known both in the Americas and the Old World. Usually such plants are treated as hybrids; in many cases they are probably just extremes of the natural variability of A. retroflexus. Putative hybrids of A. retroflexus were described from Europe as A. ×ozanonii Thellung (A. hybridus × A. retroflexus) and A. ×soproniensis Priszter & Karpáti (A. powellii × A. retroflexus) (see A. Thellung 1914–1919; S. Priszter 1958; P. Aellen 1959; F. Grüll and S. Priszter 1973). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 416. | FNA vol. 4. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Acnida | Amaranthaceae > Amaranthus > subg. Amaranthus |
Sibling taxa | ||
Synonyms | Acnida australis, Acnida alabamensis, Acnida cannabina var. australis, Acnida cuspidata | A. retroflexus var. salicifolius |
Name authority | (A. Gray) J. D. Sauer: Madroño 13: 15. (1955) | Linnaeus: Sp. Pl. 2: 991. (1753) |
Web links |
|