Amaranthus australis |
Amaranthus graecizans |
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southern amaranth, southern water-hemp |
Mediterranean amaranth, spreading pigweed, tumbleweed |
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Habit | Plants annual, pubescent in distal parts or becoming glabrescent at maturity. | |
Stems | erect, branched, stout to robust, usually 1.5–3 m (occasionally to 9 m!) × 30 cm. |
erect to ascending or decumbent, branched at or distal to base, 0.1–0.9 m. Leaves: petiole variable in length; blade lanceolate to nearly linear or rhombic-ovate to elliptic-ovate, (1.5–)2–4(–5) × 1–3 cm, base cuneate to broadly cuneate, margins entire, plane, rarely indistinctly undulate, apex subacute to obtuse or emarginate, mucronulate. |
Leaves | petiole 1/3–2/3 length of blade; blade narrowly lanceolate to narrowly ovate, 10–20 × 1–4 cm, base cuneate, margins entire, plane, apex acute or long-attenuate to acuminate. |
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Bracts | of pistillate flowers 1.5–2 mm; of staminate flowers with moderately heavy midribs, 1.5–2 mm. |
lanceolate, subspinescent, 1.5–2 mm, shorter or slightly longer than tepals. |
Inflorescences | mostly terminal, linear spikes to panicles, usually interrupted. |
axillary glomerules, green. |
Staminate flowers | tepals 5, inner tepals with moderately prominent, excurrent midribs, equal, 2–2.5 mm, apex subacute to mucronulate; stamens 5. |
intermixed with pistillate; tepals 3, equal or subequal; stamens 3. |
Pistillate flowers | tepals absent; style branches spreading; stigmas 3–5. |
tepals 3, erect, elliptic to lanceolate-elliptic, equal or subequal, 1.5–2 mm, apex short-acuminate; style branches slightly spreading; stigmas (2–)3. |
Seeds | reddish brown to dark brown, 1–1.2 mm diam., shiny. |
black, lenticular, 1–1.3(–1.6) mm diam., smooth or indistinctly punctate. |
Utricles | stramineous to brown, with 3(–5) longitudinal ridges corresponding to 3–5 style branches, elliptic or obovoid, 1.5–2.5 mm, slightly fleshy, smooth (slightly rugose in herbarium specimens). |
subglobose to broadly elliptic, 2–2.5 mm, slightly rugose, dehiscence regularly circumscissile, rarely irregularly dehiscent. |
Amaranthus australis |
Amaranthus graecizans |
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Phenology | Flowering summer–fall. | Flowering summer–fall. |
Habitat | Freshwater and brackish wetland habitats, coastal marshes, swamps, riverbanks, bayous, canals, ditches, estuaries, lakeshores, hammocks | On ballast |
Elevation | 0-100 m (0-300 ft) | 0 m (0 ft) |
Distribution |
AL; AR; FL; GA; LA; MS; NC; TN; TX; VA; e Mexico; West Indies; n South America
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NJ; native to Eurasia (Mediterranean area, s Asia); n Africa [Introduced in North America; locally introduced in Australia]
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Discussion | Plants of Amaranthus australis, a herbaceous annual, can be amazingly tall, with a single hollow main stem, up to 9 m, and the stem base can reach 30 cm in diameter. Large plants may somewhat resemble young trees of Taxodium distichum var. imbricarium, pondcypress. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In North America the name Amaranthus graecizans has been constantly misapplied to the common North American taxa A. albus and A. blitoides. Consequently, A. graecizans has been excluded from lists of North American plants. Recently, herbarium specimens (casual aliens collected in 1879 on ballast in Camden, New Jersey) of A. graecizans subsp. sylvestris were discovered (M. Costea et al. 2001b). Probably, the species disappeared in North America long ago, but, considering the long history of misidentification and confusion, there is also some chance that it may occur locally as an introduced species. Three subspecies are usually recognized within Amaranthus graecizans in the Old World: subsp. graecizans, subsp. sylvestris (Villars) Brenan, and subsp. thellungianus (Nevski) Gusev. Only subsp. sylvestris, characterized by rhombic-ovate to elliptic-ovate leaves (as compared to lanceolate to almost linear leaves in subsp. graecizans) and comparatively large seeds has so far been reported from North America. Despite its superficial similarity to Amaranthus albus and A. blitoides, A. graecizans seems to be more closely related to other Old World taxa with trimerous flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 416. | FNA vol. 4. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Acnida | Amaranthaceae > Amaranthus > subg. Albersia |
Sibling taxa | ||
Synonyms | Acnida australis, Acnida alabamensis, Acnida cannabina var. australis, Acnida cuspidata | |
Name authority | (A. Gray) J. D. Sauer: Madroño 13: 15. (1955) | Linnaeus: Sp. Pl. 2: 990. (1753) |
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