Amaranthus australis |
Amaranthus dubius |
|
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southern amaranth, southern water-hemp |
spleen amaranth |
|
Habit | Plants glabrous or sparsely pubescent in distal parts. | |
Stems | erect, branched, stout to robust, usually 1.5–3 m (occasionally to 9 m!) × 30 cm. |
erect, green, branched, 0.3–1 m. Leaves: petiole of proximal leaves equaling or longer than blade, becoming shorter distally; blade rhombic-ovate or ovate to elliptic, 3–12 × 2–8 cm, base broadly cuneate, margins entire, apex slightly acuminate to obtuse and faintly emarginate, mucronate. |
Leaves | petiole 1/3–2/3 length of blade; blade narrowly lanceolate to narrowly ovate, 10–20 × 1–4 cm, base cuneate, margins entire, plane, apex acute or long-attenuate to acuminate. |
|
Bracts | of pistillate flowers 1.5–2 mm; of staminate flowers with moderately heavy midribs, 1.5–2 mm. |
lanceolate, shorter than 2 mm, shorter than tepals, apex spinescent. |
Inflorescences | mostly terminal, linear spikes to panicles, usually interrupted. |
terminal panicles and axillary spikes; panicles erect or often drooping, green, dense, branched, leafless at least distally. |
Staminate flowers | tepals 5, inner tepals with moderately prominent, excurrent midribs, equal, 2–2.5 mm, apex subacute to mucronulate; stamens 5. |
usually clustered at tips of inflorescence branches, sometimes gathered in proximal glomerules (as in A. spinosus); tepals 5, equal or subequal; stamens 5. |
Pistillate flowers | tepals absent; style branches spreading; stigmas 3–5. |
tepals 5, oblong-spatulate to oblong, not clawed, 1.5–2 mm, apex acute, often very shortly mucronate; style branches strongly spreading, shorter than body of fruit; stigmas 3. |
Seeds | reddish brown to dark brown, 1–1.2 mm diam., shiny. |
dark reddish brown to black, subglobose or lenticular, 0.8–1 mm diam., shiny, smooth. |
Utricles | stramineous to brown, with 3(–5) longitudinal ridges corresponding to 3–5 style branches, elliptic or obovoid, 1.5–2.5 mm, slightly fleshy, smooth (slightly rugose in herbarium specimens). |
ovoid or subglobose, 1.5–2 mm, slightly shorter than tepals, smooth to irregularly wrinkled, dehiscence regularly circumscissile. |
2n | = 64. |
|
Amaranthus australis |
Amaranthus dubius |
|
Phenology | Flowering summer–fall. | Flowering summer–fall in tropics, various seasons in subtropics. |
Habitat | Freshwater and brackish wetland habitats, coastal marshes, swamps, riverbanks, bayous, canals, ditches, estuaries, lakeshores, hammocks | Waste places, disturbed habitats |
Elevation | 0-100 m (0-300 ft) | 0-100 m (0-300 ft) |
Distribution |
AL; AR; FL; GA; LA; MS; NC; TN; TX; VA; e Mexico; West Indies; n South America
|
FL; South America; West Indies [Introduced in North America; introduced and locally naturalized Europe, Asia, Africa] |
Discussion | Plants of Amaranthus australis, a herbaceous annual, can be amazingly tall, with a single hollow main stem, up to 9 m, and the stem base can reach 30 cm in diameter. Large plants may somewhat resemble young trees of Taxodium distichum var. imbricarium, pondcypress. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Amaranthus dubius, a morphologically deviant allopolyploid, is very close genetically to both A. spinosus (sect. Centrusa) and members of sect. Amaranthus. This species most probably originated as a result of ancient hybridization between A. spinosus and either A. hybridus or A. quitensis (W. F. Grant 1959; T. N. Khoshoo and M. Pal 1972; M. Pal and T. N. Khoshoo 1965; J. D. Sauer 1967b; V. Srivastava et al. 1977). Amaranthus nothosect. Dubia Mosyakin & K. R. Robertson (A. sect. Amaranthus × A. sect. Centrusa), was proposed to accommodate A. dubius (S. L. Mosyakin and K. R. Robertson 1996). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 416. | FNA vol. 4, p. 425. |
Parent taxa | Amaranthaceae > Amaranthus > subg. Acnida > sect. Acnida | Amaranthaceae > Amaranthus > subg. Amaranthus |
Sibling taxa | ||
Synonyms | Acnida australis, Acnida alabamensis, Acnida cannabina var. australis, Acnida cuspidata | |
Name authority | (A. Gray) J. D. Sauer: Madroño 13: 15. (1955) | Martius ex Thellung: Fl. Adv. Montpellier 38: 203. (1912) |
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