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foxtail, meadow-foxtail

alpine foxtail, alpine meadow-foxtail, boreal foxtail, foxtail, vulpin alpin, vulpin boreale

Habit Plants annual or perennial; sometimes cespitose, sometimes shortly rhizomatous. Plants perennial; shortly rhizomatous.
Culms

5-110 cm, clumped or solitary, erect or decumbent, occasionally cormlike at the base;

nodes glabrous.

(6)10-80 cm, erect or decumbent.

Leaves

inserted mostly on the lower 1/2 of the culms;

sheaths open, upper sheaths sometimes inflated;

auricles absent;

ligules 0.6-6.5 mm, truncate to acute, membranous, puberulent or glabrous, entire to lacerate;

blades 0.7-12 mm wide, flat or involute, glabrous or scabrous, blades of uppermost leaves sometimes short or absent.

Panicles

1-5 cm long, 8-14 mm wide.

Inflorescences

terminal panicles, spikelike, capitate to cylindrical;

branches usually shorter than 5 mm, lower branches sometimes to 2 cm;

disarticulation below the glumes.

Spikelets

1.8-7 mm, pedicellate, strongly laterally compressed, oval in outline, with 1 floret;

rachillas not prolonged beyond the base of the floret.

Glumes

equaling or exceeding the florets, membranous or coriaceous, free or connate in at least the lower 1/2, narrowing from above midlength, 3-veined, keeled, keels ciliate, at least basally, apices obtuse to acute or shortly awned;

calluses blunt, glabrous;

lemmas membranous, margins often connate in the lower 1/2, keeled, indistinctly 3-5-veined, apices truncate to acute, awned dorsally from just above the base to about midlength, geniculate or straight;

paleas absent or greatly reduced;

lodicules absent;

anthers 3, 0.3-4.1 mm;

ovaries glabrous;

styles fused, with 2 branches.

3-5 mm, connate in the lower 1/8, membranous, densely pilose throughout, keels not winged, ciliate, apices acute and parallel;

lemmas 2.5-4.5 mm, connate in the lower 1/2 - 2/3, glabrous proximally, finely pubescent distally, apices usually obtuse, occasionally truncate, awns 2-6(8) mm, geniculate, exceeding the lemmas by 0-5 mm;

anthers 2.3-3 mm, yellow.

Caryopses

shorter than the lemmas, concealed at maturity, glabrous;

hila short, x = 7.

0.7-2 mm.

Ligules

1-2 mm, truncate;

blades 4-22 cm long, 2.5-7 mm wide;

upper sheaths inflated.

2n

= 98, 100, 105, 112, 117, 119, ca. 120.

Alopecurus

Alopecurus magellanicus

Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
[BONAP county map]
from FNA
AK; CO; ID; MT; UT; WY; AB; BC; MB; NF; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Alopecurus is a genus of 36 species that grow primarily in open, mesic habitats, and are native to the northern temperate zone and South America. Four species are native to the Flora region, four were introduced and have become established, and two were introduced and are not known to persist. Some species, including some native to the Flora region, have been introduced as pasture grasses outside of their native ranges. Of these, only A. pratensis has become widely naturalized.

Some species of Alopecurus can appear similar to Phleum, which has truncate glumes that are abruptly awned or mucronate, lemmas without awns or keels, and well-developed paleas; Alopecurus has glumes that are obtuse to acute and gradually awned or unawned, lemmas with both awns and keels, and paleas that are absent or greatly reduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Alopecurus magellanicus has an arctic-alpine to subalpine circumpolar distribution, but it has not been found in Scandinavia or Iceland. It grows primarily in wet soils in tundra, meadows, along streams, shorelines, gravelbars, and floodplains, and occasionally in somewhat drier forest openings, in fine or silty to stony soils or moss. It is sometimes co-dominant with Dupontia fisheri in the arctic and subarctic portion of its range. The anthocyanic tint of the plant as a whole greatly increases to the north.

In the past, this species has been called Alopecurus alpinus, the name being attributed to Smith. Dogan (1999) pointed out that the name had first been used by Villars for a different species, which meant that this species had to have another name; Dogan used A. borealis Trin., listing A. magellanicus as a synonym but, because A. magellanicus was published first, it has priority and is therefore the correct name.

The morphological variability in Alopecurus magellanicus has prompted recognition of several segregate taxa, A. stejnegeri Vasey and A. occidentalis Scribn. & Tweedy being two of the more conspicuous extremes. The former are small plants occurring on enriched sites in the arctic, usually around seabird or seal colonies where high nutrient levels produce lush vegetative growth; the latter refers to tall-stemmed plants found in the Rocky Mountains. Because such plants are simply extremes in a continuum of variation, they do not merit taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Glume keels winged; glumes glabrous, pubescent over the veins.
→ 2
2. Glumes 4.5-7.5 mm long, connate in the lower 1/2, the apices acute, convergent to parallel; lemma apices acute
A. myosuroides
2. Glumes 3-4.5 mm long, connate in the lower 1/2 - 4/5, the apices obtuse, mucronate, divergent; lemma apices truncate
A. creticus
1. Glume keels not winged; glumes usually sparsely to densely pubescent, sometimes glabrous.
→ 3
3. Plants annual, without rhizomes, not rooting at the lower nodes; blades 1-16 cm long, 0.9-4 mm wide; culms 5-50 cm tall.
→ 4
4. Glumes 5-6.4 mm long, coriaceous and dilated in the lower 1/2; glume and lemma apices acute to acuminate; anthers about 3 mm long
A. rendlei
4. Glumes 2.1-5 mm long, membranous, not dilated below; glume and lemma apices obtuse; anthers 0.3-1.8 mm long.
→ 5
5. Upper sheaths conspicuously inflated; glumes 3-5 mm long; lemmas 3-5 mm long, awns exceeding the lemmas by 3-6 mm; panicles 5.5-13 mm wide, excluding the awns
A. saccatus
5. Upper sheaths not or only slightly inflated; glumes 2.1-3.1 mm long; lemmas 1.9-2.7 mm long, awns exceeding the lemmas by 1.6-4 mm; panicles 3-6 mm wide, excluding the awns
A. carolinianus
3. Plants perennial, often rhizomatous, sometimes rooting at the lower nodes; blades 2-40 cm long, 1-12 mm wide; culms 5-110 cm tall.
→ 6
6. Glumes 1.8-3.7 mm long, the apices obtuse; anthers 0.5-2.2 mm long.
→ 7
7. Awns geniculate, exceeding the lemmas by 1.2-4 mm; anthers (0.9)1.4-2.2 mm long
A. geniculates
7. Awns straight, not exceeding the lemmas or exceeding them by less than 2.5 mm; anthers 0.5-1.2 mm long
A. aequalis
6. Glumes 3-6 mm long, the apices acute; anthers 2-4 mm long.
→ 8
8. Glume margins connate in the lower 1/8; glumes densely pilose throughout
A. magellanicus
8. Glume margins connate in the lower 1/5 – 1/3; glumes with long hairs mainly restricted to the veins.
→ 9
9. Lemma apices acute; glume apices parallel or convergent
A. pratensis
9. Lemma apices obtuse to truncate; glume apices divergent
A. arundinaceus
Source FNA vol. 24, p. 780. Treatment author: William J. Crins;. FNA vol. 24, p. 782. Treatment author: William J. Crins.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae Poaceae > subfam. Pooideae > tribe Poeae > Alopecurus
Sibling taxa
A. aequalis, A. arundinaceus, A. carolinianus, A. creticus, A. geniculatus, A. myosuroides, A. pratensis, A. rendlei, A. saccatus
Subordinate taxa
A. aequalis, A. arundinaceus, A. carolinianus, A. creticus, A. geniculates, A. magellanicus, A. myosuroides, A. pratensis, A. rendlei, A. saccatus
Synonyms A. occidentalis, A. borealis, A. alpinus subsp. glaucus, A. alpinus
Name authority L. Lam.
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