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agrostide stolonifere, carpet bentgrass, creeping bent, creeping bentgrass, fiorin, redtop, spreading bent

bentgrass, dryland bentgrass, dryland browntop, highland bent, highland bentgrass

Habit Plants perennial; stoloniferous, stolons 5-100+ cm, rooting at the nodes, often forming a dense mat, without rhizomes. Plants perennial; loosely cespitose, rhizomatous, rhizomes to 10(40) cm, covered with inflated scales.
Culms

(8)15-60 cm, erect from a geniculate base, sometimes rooting at the lower nodes, with (2)4-7 nodes.

30-80 cm, erect or geniculate, with up to 10 nodes, basal internodes devoid of leaf bases after anthesis in dry habitats, shoots proliferating from the upper nodes, especially of the innovations, later in the season.

Leaves

mostly cauline;

sheaths smooth;

ligules longer than wide, dorsal surfaces usually scabrous, rarely smooth, apices usually rounded, acute to truncate, erose to lacerate, basal ligules 0.7-4 mm, upper ligules 3-7.5 mm;

blades 2-10 cm long, 2-6 mm wide, flat.

mostly cauline;

sheaths smooth;

ligules 0.5-3 mm, shorter than wide, dorsal surfaces smooth or scabridulous, apices truncate to rounded, ciliolate-erose;

blades 4-10 cm long, 1-3 mm wide, flat.

Panicles

(3)4-20 cm long, less than 1/2 the length of the culm, 0.5-3(6) cm wide, narrowly contracted, dense, oblong to lanceolate, exserted from the sheaths at maturity, lowest node with 1-7 branches;

branches scabrous, ascending to appressed, except briefly spreading during anthesis, usually some branches at each node spikelet-bearing to the base, lower branches 2-6 cm;

pedicels 0.3-3.3 mm.

10-30 cm long, less than 1/2 the length of the culm, 3-8 cm wide, loosely ovate, somewhat lax, somewhat contracted and linear-lanceolate after anthesis, lowest node with (1)2-7 branches;

branches spreading to ascending, sparsely scabridulous, branching above the midpoint, spikelets usually confined to the distal 1/3 and in discrete clusters at the branch tips, lower branches 3-9 cm;

pedicels 0.6-2.3(3) mm, usually shorter than the spikelets, adjacent pedicels not divergent.

Spikelets

lanceolate, green and slightly to strongly suffused with purple.

lanceolate, yellowish green to stramineous or brownish, slightly to strongly suffused with purple.

Glumes

subequal to unequal, 1.6-3 mm, lanceolate, 1-veined, sometimes scabridulous distally, at least on the midvein, acute to acuminate or apiculate;

callus hairs to 0.5 mm, sparse;

lemmas 1.4-2 mm, opaque to translucent, smooth, 5-veined, veins obscure or prominent distally, apices acute to obtuse, entire or the veins excurrent to about 0.1 mm, usually unawned, rarely with a subapical straight awn to about 1 mm;

paleas 0.7-1.4 mm, veins visible;

anthers 3, 0.9-1.4 mm.

subequal, 2-3 mm, lanceolate, 1-veined, acute or acuminate;

lower glumes scabrous over the midvein, at least distally;

upper glumes smooth or scabrous to scabridulous over the midvein distally;

calluses abundantly hairy, hairs to 0.3(0.6) mm;

lemmas 1.3-1.9 mm, occasionally with hairs on the lower 1/2, translucent, (3)5-veined, veins prominent distally, apices usually truncate to obtuse, sometimes acute, entire or the lateral veins excurrent to 0.6 mm, awned or unawned, usually mixed in the inflorescence, terminal spikelets usually awned, in some plants all unawned, awns to 5 mm, awns arising from the lower 1/3 or occasionally as a minute bristle from above the center;

paleas 0.6-1.1 mm, 1/2-2/3 as long as the lemma;

anthers 3, 1-1.5 mm.

Caryopses

0.9-1.3 mm;

endosperm solid.

about 1 mm.

2n

= 28, 35,42.

= 28, 42.

Agrostis stolonifera

Agrostis castellana

Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; Greenland
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[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Agrostis stolonifera grows in areas that are often temporarily flooded, such as lakesides, marshes, salt marshes, lawns, and damp fields, as well as moist meadows, forest openings, and along streams. It will also colonize disturbed sites such as ditches, clearcuts, and overgrazed pastures. Its North American range extends from the subarctic into Mexico, mostly at low to middle elevations.

Agrostis stolonifera has been confused with A. gigantea (see previous). It is considered to be Eurasian, but some northern salt marsh and lakeside populations may be native. Agrostis stolonifera is also similar to A. castellana (p. 639); it differs in having longer, acute to truncate ligules that are longer than wide, and in possessing extensive stolons. The names A. palustris Huds. and A. maritima Lam. have been applied to plants with longer stolons; all forms intergrade. A hybrid between A. stolonifera and Polypogon monspeliensis, xAgropogon lutosus (p. 668), has been found in the Flora region. It differs from A. stolonifera in having awned glumes and lemmas. Agrostis stolonifera readily hybridizes with A. vinealis (see below), the hybrids being somewhat intermediate between the two parents.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Agrostis castellana is native to southern Europe. It was introduced to North America in the 1930s for use in lawns and golf greens, under the name Agrostis tenuis 'Highland'; commercial samples of 'Highland' often contain A. capillaris (p. 639). Escaped plants were collected at least as early as the 1950s, but were not recognized as belonging to A. castellana until the 1990s, when several collections were identified as such in Oregon. Recorded habitats have ranged from sunny gravel roadsides to moist ground alongside cranberry bogs, at elevations from near sea level to over 600 m.

In view of its extensive commercial use for over 70 years and its drought tolerance, it is likely that it is more widespread than shown.

Agrostis castellana belongs to a Eurasian group that includes A. gigantea (p. 642), A. stolonifera (p. 641), and A. capillaris (635). It differs from A. gigantea and A. stolonifera in having shorter, truncate ligules about as short as wide, and in not possessing extensive rhizomes and stolons. It differs from A. capillaris in having clustered rather than diffuse spikelets, more abundant rhizomes, somewhat constricted panicle branches after anthesis, abundantly hairy calluses with hairs up to 0.3(0.6) mm long, and lemmas that are sometimes dorsally pubescent. It also tends to flower somewhat later than A. capillaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 641. FNA vol. 24, p. 639.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Agrostis Poaceae > subfam. Pooideae > tribe Poeae > Agrostis
Sibling taxa
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. exarata, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. tolucensis, A. variabilis, A. vinealis
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. clavata, A. densiflora, A. elliottiana, A. exarata, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
Synonyms A. stolonifera var. palustris, A. stolonifera var. compacta, A. palustris, A. maritima, A. alba var. stolonifera, A. alba var. palustris, A. alba forma aristigera
Name authority L Boiss. & Reut.
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