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bentgrass, spike bent, spike bent grass

agrostide scabre, foin fou, hair bentgrass, rough bent, rough bent grass, ticklegrass, winter bentgrass

Habit Plants perennial; usually cespitose, sometimes rhizomatous, not stoloniferous. Plants perennial or annual; cespitose, not rhizomatous or stoloniferous.
Culms

8-100 cm, erect or decumbent at the base, sometimes rooting at the lower nodes, with (2)3-6(8) nodes.

(7.5) 15-90 cm, erect, nodes usually 1-3.

Leaves

mostly cauline;

sheaths smooth or slightly scabrous;

ligules (1)1.7-8(11.2) mm, dorsal surfaces scabrous, apices truncate to obtuse, lacerate to erose;

blades 4-15 cm long, 2-7 mm wide, flat.

mostly basal, basal leaves usually persistent;

sheaths usually smooth, sometimes scabridulous;

ligules 0.7-5 mm, dorsal surfaces scabrous, apices usually rounded, sometimes truncate or acute, erose-ciliolate, sometimes lacerate;

blades 4-14 cm long, 1-2 mm wide, basal blades mostly involute;

cauline blades mostly flat.

Panicles

(3)5-30 cm long, 0.5-4 cm wide, contracted, spikelike, oblong, or lanceolate, usually dense, rarely more open, sometimes interrupted near the base, bases usually exserted, rarely enclosed by the upper sheaths at maturity, lowest node with 1-5 branches;

branches scabrous, ascending to appressed, spikelet-bearing to or near the base, usually hidden by the spikelets, spikelets crowded, lower branches 1-2(4) cm;

pedicels 0.2-4.3 mm.

(4)8-25(50) cm long, 0.5-20 cm wide, broadly ovate, often nearly as wide as long, diffuse, the whole panicle often detaching at the base at maturity, forming a tumbleweed, exserted from the upper sheaths, lowest node with (1)2-7(12) branches;

branches scabrous, capillary, flexible, wide-spreading, readily visible, branching beyond midlength, spikelets somewhat distant, not crowded, lower branches 4-12 cm;

pedicels 0.4-9.6 mm.

Spikelets

lanceolate to narrowly ovate, greenish to purplish.

lanceolate, greenish purple, frequently purple at maturity.

Glumes

subequal to equal, 1.5-3.5 mm, scabrous on the midvein and sometimes on the back, 1(3)-veined, acute, elongate-acuminate, with an awnlike tip to 1 mm;

callus hairs to 0.3 mm, sparse to abundant;

lemmas 1.2-2.2 mm, smooth, translucent to opaque, 5-veined, veins prominent distally or obscure throughout, apices acute, entire or toothed, teeth no more than 0.12 mm, unawned or awned from above midlength, awns to 3.5 mm, straight or geniculate;

paleas absent or to 0.5 mm;

anthers 3, 0.3-0.6 mm.

unequal, 1.8-3.4 mm, lanceolate, 1-veined, keels scabrous at least towards the apices, apices acuminate;

callus hairs to 0.2 mm, sparse;

lemmas 1.4-2 mm, scabrous to scabridulous or smooth, translucent to opaque, 5-veined, veins prominent, apices acute to obtuse, usually entire, sometimes minutely toothed, unawned or awned from below midlength, awns 0.2-3 mm, exceeding the lemma apices by up to 2.5 mm, geniculate or straight, persistent;

paleas absent or to 0.2 mm;

anthers 3, 0.4-0.8 mm, usually shed at anthesis.

Caryopses

0.9-1.2 mm;

endosperm solid or soft.

0.9-1.4 mm;

endosperm liquid.

2n

= 28, 42, 56.

= 42.

Agrostis exarata

Agrostis scabra

Distribution
from FNA
AK; AZ; CA; CO; ID; KY; MT; NE; NM; NV; OK; OR; SD; TX; UT; VT; WA; WY; AB; BC; NT; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DE; FL; GA; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MN; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Agrostis exarata is common and widely distributed in western North America, usually growing in moist ground in open woodlands, river valleys, tidal marshes, and swamp and lake margins; it also grows in dry habitats such as grasslands and shrublands. It extends from Alaska into Mexico, and is also found in Kamchatka and the Kuril Islands. Eastern North American records probably reflect introductions. It readily colonizes roadsides and bare soil, and exhibits ecological and developmental flexibility. Agrostis exarata is recognized here as a single, variable species that includes what others have treated as distinct species or varieties. Cytotaxonomic study might clarify the basis of the observed variation. Agrostis exarata appears to be related to A. densiflora (see below).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Agrostis scabra grows in a wide variety of habitats, including grasslands, meadows, shrublands, woodlands, marshes, and stream and lake margins, as well as disturbed sites such as roadsides, ditches, and abandoned pastures. It occurs throughout much of the Flora region, but is not common in the Canadian high arctic or the southeastern United States. It extends south into Mexico; it is also native to the Pacific coast from Kamchatka to Japan and Korea, and has been introduced elsewhere.

Plants in the Agrostis scabra aggregate are variable. Awned and unawned plants often occur together, the difference presumably being caused by a single gene. At least three groups may be distinguished within the species as treated here: widespread, lowland, rather weedy plants capable of producing very large panicles that have been introduced into the southern United States; smaller, short-leaved, slow-growing plants of rocks and screes, which are widespread in the Rockies, the Appalachians, and much of Alaska, Canada, and Greenland; and luxuriant, broad-leaved plants that are characteristically found in sheltered, frost-free canyons of the southwestern United States. The second group has sometimes been called A. scabra var. geminata (Trin.) Swallen or A. geminata Trin.

Tercek et al. (2003) found that annual forms of Agrostis scabra with inflated upper sheaths and open panicles that were collected around hot springs in western North America were molecularly, and in some respects morphologically, more similar to plants identified as hot spring endemics such as A. rossiae and A. pauzbetica Prob., than they were to neighboring perennial plants of A. scabra that did not have inflated leaf sheaths. They differed, however, in having open, rather than contracted, panicles.

Agrostis scabra is often confused with a number of other species; for comparisons, see under the appropriate species description: A. mertensii (p. 644), A. clavata (see previous), A. hyemalis (see next), A. perennans (see this page), and A. idahoensis (p. 649).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 651. FNA vol. 24, p. 646.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Agrostis Poaceae > subfam. Pooideae > tribe Poeae > Agrostis
Sibling taxa
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. exarata, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
Synonyms A. longiligula var. australis, A. longiligula, A. exarata var. purpurascens, A. exarata var. pacifica, A. exarata var. monolepis, A. exarata var. minor, A. exarata subsp. minor, A. ampla, A. alaskana A. scabra forma tuckermanii, A. scabra var. septentrionalis, A. scabra. var. geminata, A. hyemalis var. tenuis, A. hyemalis var. scabra, A. hyemalis var. geminata, A. geminata
Name authority Trin. Willd.
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