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bentgrass, spike bent, spike bent grass

agrostide d'hiver, tickle grass, winter bent, winter bentgrass

Habit Plants perennial; usually cespitose, sometimes rhizomatous, not stoloniferous. Plants perennials or facultative annuals; cespitose, not rhizomatous or stoloniferous.
Culms

8-100 cm, erect or decumbent at the base, sometimes rooting at the lower nodes, with (2)3-6(8) nodes.

15-82 cm, erect, with (3)4-7 nodes.

Leaves

mostly cauline;

sheaths smooth or slightly scabrous;

ligules (1)1.7-8(11.2) mm, dorsal surfaces scabrous, apices truncate to obtuse, lacerate to erose;

blades 4-15 cm long, 2-7 mm wide, flat.

cauline and basal;

sheaths smooth;

ligules (0.7) 1.2-4 mm, dorsal surfaces scabrous, apices usually rounded to truncate, sometimes acute, lacerate-erose;

blades 3-10 cm long, 1-2 mm wide, flat, becoming involute, or folded.

Panicles

(3)5-30 cm long, 0.5-4 cm wide, contracted, spikelike, oblong, or lanceolate, usually dense, rarely more open, sometimes interrupted near the base, bases usually exserted, rarely enclosed by the upper sheaths at maturity, lowest node with 1-5 branches;

branches scabrous, ascending to appressed, spikelet-bearing to or near the base, usually hidden by the spikelets, spikelets crowded, lower branches 1-2(4) cm;

pedicels 0.2-4.3 mm.

(5)10-25(36) cm long, (3)4-24 cm wide, broadly ovate, often nearly as wide as long, diffuse, the whole panicle often detaching at the base at maturity, forming a tumbleweed, bases often enclosed by the upper sheaths, lowest node with (3)5-11 branches;

branches scabrous, capillary, flexible, wide-spreading, branching in the distal 1/4, spikelets strongly clustered at the branch tips, lower branches 5-15 cm;

pedicels 0.1-2.5(3.5) mm.

Spikelets

lanceolate to narrowly ovate, greenish to purplish.

ovate to narrowly ovate, greenish or purplish.

Glumes

subequal to equal, 1.5-3.5 mm, scabrous on the midvein and sometimes on the back, 1(3)-veined, acute, elongate-acuminate, with an awnlike tip to 1 mm;

callus hairs to 0.3 mm, sparse to abundant;

lemmas 1.2-2.2 mm, smooth, translucent to opaque, 5-veined, veins prominent distally or obscure throughout, apices acute, entire or toothed, teeth no more than 0.12 mm, unawned or awned from above midlength, awns to 3.5 mm, straight or geniculate;

paleas absent or to 0.5 mm;

anthers 3, 0.3-0.6 mm.

subequal, 1-2 mm, 1-veined, keeled, keels scabrous, sometimes the body also scabrous towards the apices, acute to acuminate;

callus hairs to 0.2 mm, sparse;

lemmas 0.8-1.2 mm, scabridulous, translucent to opaque, 5-veined, veins obscure or prominent distally, apices usually obtuse, sometimes acute, entire, unawned;

paleas absent, or to 0.2 mm and thin;

anthers 3, 0.2-0.5 mm.

Caryopses

0.9-1.2 mm;

endosperm solid or soft.

0.7-1 mm.

2n

= 28, 42, 56.

= 28.

Agrostis exarata

Agrostis hyemalis

Distribution
from FNA
AK; AZ; CA; CO; ID; KY; MT; NE; NM; NV; OK; OR; SD; TX; UT; VT; WA; WY; AB; BC; NT; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; PR; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Agrostis exarata is common and widely distributed in western North America, usually growing in moist ground in open woodlands, river valleys, tidal marshes, and swamp and lake margins; it also grows in dry habitats such as grasslands and shrublands. It extends from Alaska into Mexico, and is also found in Kamchatka and the Kuril Islands. Eastern North American records probably reflect introductions. It readily colonizes roadsides and bare soil, and exhibits ecological and developmental flexibility. Agrostis exarata is recognized here as a single, variable species that includes what others have treated as distinct species or varieties. Cytotaxonomic study might clarify the basis of the observed variation. Agrostis exarata appears to be related to A. densiflora (see below).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Agrostis hyemalis is most abundant along roadsides and in open pastures, scrub, and rocky areas. It is centered in the southeastern United States; historically it extended north to coastal Maine, where it may be extinct, west to Wisconsin and Texas, and south into the Caribbean, Mexico, and Ecuador. Records from further north and west in North America are confused; many reflect the former inclusion of the generally more northern A. scabra in A. hyemalis. Agrostis hyemalis differs from A. scabra in its smaller spikelets and anthers, more conspicuous culm leaves, and more clustered spikelets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 651. FNA vol. 24, p. 647.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Agrostis Poaceae > subfam. Pooideae > tribe Poeae > Agrostis
Sibling taxa
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. exarata, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
Synonyms A. longiligula var. australis, A. longiligula, A. exarata var. purpurascens, A. exarata var. pacifica, A. exarata var. monolepis, A. exarata var. minor, A. exarata subsp. minor, A. ampla, A. alaskana A. hiemalis
Name authority Trin. (Walter) Britton, Sterns &
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