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California bent, California bent grass

agrostide blanche, black bent, creeping bentgras, redtop, redtop bentgrass

Habit Plants perennial; not rhizomatous or stoloniferous. Plants perennial; rhizomatous, rhizomes to 25 cm, not stoloniferous.
Culms

9-85 cm, erect, sometimes decumbent at the base, sometimes rooting at the lower nodes, usually with 4-7 nodes.

20-120 cm, erect, sometimes geniculate at the base, sometimes rooting at the lower nodes, with 4-7 nodes.

Leaves

basal and cauline;

sheaths smooth or scabrous;

ligules 1-4.8(7.5) mm, dorsal surfaces scabrous, apices truncate to obtuse, erose-lacerate, sometimes ciliolate;

blades 2-12 cm long, 2-10 mm wide, upper blades broader than those below, flat.

mostly cauline, sheaths smooth or sparsely scabridulous;

ligules longer than wide, dorsal surfaces usually scabrous, sometimes smooth, apices rounded to truncate, erose to lacerate, basal ligules 1-4.5 mm, upper ligules 2-7 mm;

blades 4-10 cm long, 3-8 mm wide, flat.

Panicles

2-10 cm long, 0.5-2 cm wide, narrow and spikelike to dense, lobed-lanceolate, bases exserted or enclosed by the upper sheaths at maturity;

branches to 1.5 cm, scabrous, appressed, branching at the base and mostly hidden by the spikelets;

pedicels about 0.4-3 mm.

8-25(30) cm long, less than 1/2 the length of the culm, (1.5)3-15 cm wide, erect, open, broadly ovate, exserted from the upper sheaths at maturity, lowest node with (1)3-8 branches;

branches scabrous, spreading during and after anthesis, usually some branches spikelet-bearing to the base, lower branches 4-9 cm, usually with many shorter secondary branches resulting in crowding of the spikelets, spikelets restricted to the distal 1/2 of the branches and not crowded in shade plants;

pedicels 0.3-3.4(4.2) mm.

Spikelets

lanceolate to narrowly ovate, yellowish, sometimes tinged with purple, or greenish purple.

narrowly ovate to lanceolate, green and slightly to strongly suffused with purple.

Glumes

generally equal, 2-3.3 mm, 1-veined, densely scabrous, aculeolate on the veins, narrowly acute to acuminate or somewhat mucronate, mucros to about 0.5 mm;

callus hairs to 0.3 mm, usually dense;

lemmas 1.5-2.1 mm, smooth or scabridulous, translucent, (3)5-veined, veins prominent to obscure, sometimes prominent only distally, apices acute to obtuse, veins usually extended as teeth up to 0.3 mm, unawned or awned from above midlength, awns to 3.5 mm, straight, readily deciduous;

paleas 0.3-0.7 mm, thin;

anthers 3, 0.5-2 mm.

subequal, 1.7-3.2 mm, lanceolate, 1-veined, acute to apiculate;

lower glumes scabrous on the distal 1/2 of the midvein;

upper glumes scabridulous on the distal 1/2 of the midvein;

callus hairs to 0.5 mm, sparse;

lemmas 1.5-2.2 mm, opaque to translucent, smooth, 3-5-veined, veins usually obscure, sometimes prominent throughout or distally, often excurrent to 0.2 mm, apices usually acute, sometimes obtuse or truncate, usually unawned, rarely with a 0.4-1.5(3) mm straight awn arising from near the apices to near the base;

paleas 0.7-1.4 mm, about Vi the length of the lemmas, veins visible;

anthers 3, 1-1.4 mm.

Caryopses

1-1.5 mm;

endosperm solid.

1-1.5 mm;

endosperm solid.

2n

= 42.

= 42.

Agrostis densiflora

Agrostis gigantea

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; PE; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Agrostis densiflora is endemic to coastal Oregon and California. It grows in sandy soils, on cliffs, and in scrublands. It appears to be related to A. exarata (see previous), and hybridizes with A. blasdalei (p. 656).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Agrostis gigantea grows in fields, roadsides, ditches, and other disturbed habitats, mostly at lower elevations. It is a serious agricultural weed, as well as a valuable soil stabilizer. In the Flora region, its range extends from the subarctic to Mexico; it is considered to be native to Eurasia. It is more heat tolerant than most species of Agrostis.

Agrostis gigantea has been confused with A. stolonifera (see next), from which it differs in having rhizomes and a more open panicle. Agrostis stolonifera has elongated leafy stolons, mainly all above the surface, that root at the nodes, and the panicles are condensed and often less strongly pigmented than in A. gigantea. Its distribution tends to be more northern and coastal where ditches and pond margins are common habitats, and its stolons enable it to form loose mats. Agrostis gigantea is ecologically adapted to a more extreme climate—hot summers/cold winters and drought—than A. stolonifera. It is also similar to A. capillaris (p. 639) and A. castellana (p. 639); it differs from both in its longer ligules, from A. capillaris in its less open panicles with spikelets near the base of the branches, and from A. castellana in being more extensively rhizomatous.

When Agrostis gigantea grows in damp hollows under trees it becomes more like A. stolonifera, particularly when the inflorescence is young, not expanded, and pale. If the rootstock is not collected, identification is a major problem.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 651. FNA vol. 24, p. 641.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Agrostis Poaceae > subfam. Pooideae > tribe Poeae > Agrostis
Sibling taxa
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. elliottiana, A. exarata, A. gigantea, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
A. anadyrensis, A. blasdalei, A. canina, A. capillaris, A. castellana, A. clavata, A. densiflora, A. elliottiana, A. exarata, A. hallii, A. hendersonii, A. hooveri, A. howellii, A. hyemalis, A. idahoensis, A. mertensii, A. microphylla, A. nebulosa, A. oregonensis, A. pallens, A. perennans, A. rossiae, A. scabra, A. stolonifera, A. tolucensis, A. variabilis, A. vinealis
Synonyms A. clivicola var. punta-reyensis, A. californica A. stolonifera var. major, A. gigantea var. ramosa, A. gigantea var. dispar, A. alba var. major
Name authority Vasey Roth
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