The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

incised agrimony, incised groovebur

Habit Herbs, 3–11 dm. Herbs, perennial (annual or biennial in Poteridium) [shrubs or trees]; unarmed (hypanthia armed in Acaena).
Roots

tubers fusiform, thickened.

Stems

with glistening sessile-glandular hairs and ± matted-pubescent to villous and hirsute (hairs scattered, erect, stiff, 2–3 mm).

Leaves

mid cauline stipules ± falcate, margins dentate;

major leaflets 3–15 (mid cauline 9), minor 1–3 pairs;

major leaflet blades ± obovate, terminal largest, largest of these 2.3–4.1 × 1–1.7 cm, margins incised, apex obtuse to acute, abaxial surfaces glistening with sessile-glandular hairs and pubescent to villous and sparsely hirsute (hairs stiff, 2 mm).

alternate, odd-pinnately compound;

stipules persistent (absent in Acaena), adnate to petiole (free in Poterium);

venation pinnate.

Inflorescences

axes often with glistening sessile-glandular hairs and pubescent to villous and hirsute (hairs stiff, erect, 2 mm).

Flowers

usually ± alternate.

perianth and androecium perigynous;

epicalyx bractlets absent;

hypanthium hemispheric, obconic, ovoid, urceolate, top-shaped, ellipsoid, nearly orbicular, or obtriangular;

torus absent;

carpels 1 or 2(or 3), rarely more, styles terminal, distinct;

ovule 1, apical.

Fruits

achenes, enclosed within enlarged, often hardened, sometimes armed hypanthia;

styles deciduous, not elongate.

Fruiting

hypanthia hemispheric to turbinate, 1.5–2.7 × 1.8–3.3 mm, obscurely sulcate, hooked bristles in 3–4 circumferential rows, proximal row spreading 45–90°, glistening with sessile-glandular hairs (often yellow).

Agrimonia incisa

Rosaceae tribe Agrimonieae

Phenology Flowering late Jul–Nov.
Habitat Dry to moist, longleaf pine-oak woods, oak-hickory slopes, roadsides, sand or shell maritime thickets
Elevation 0–200 m (0–700 ft)
Distribution
from FNA
AL; FL; GA; LA; MO; MS; SC; TX
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; South America; West Indies (Hispaniola); Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands (New Zealand); Australia
Discussion

Populations of Agrimonia incisa are infrequent and usually widely scattered within the limited range (15–20 currently known). The plants are sometimes abundant where they occur.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 12, species ca. 270 (5 genera, 17 species in the flora).

The base chromosome number for Agrimonieae is x = 7. Acaena and Sanguisorba are host to Phragmidium rusts. The tribal name Agrimonieae has priority over Sanguisorbeae, used by, among others, D. Potter et al. (2007). Agrimonieae also includes the genera Aremonia Necker ex Nestler (Europe), Cliffortia Linnaeus, Hagenia J. F. Gmelin and Leucosidea Ecklon & Zeyher (Africa), Margyricarpus Ruiz & Pavón and Polylepis Ruiz & Pavón (South America), and Spenceria Trimen (Asia) (Potter et al.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems creeping or suberect; hypanthia spiny.
Acaena
1. Stems ascending to erect; hypanthia not spiny (rim bristly in fruit in Agrimonia)
→ 2
2. Inflorescences racemes (simple or compound); leaf blades with minor leaflet pairs between major pairs; petals 5 (yellow); hypanthial rim with 2–5 rows of bristles.
Agrimonia
2. Inflorescences spikes (cylindric, globose, or headlike); leaf blades without minor leaflet pairs between major pairs; petals 0; hypanthial rim without rows of bristles
→ 3
3. Leaflet margins pectinately pinnatisect.
Poteridium
3. Leaflet margins crenate, serrate, or incised
→ 4
4. Spikes headlike; flowers bisexual or pistillate (plants gynomonoecious); sepals distinct.
Poterium
4. Spikes ellipsoid to cylindric; flowers bisexual; sepals basally connate.
Sanguisorba
Source FNA vol. 9, p. 314. FNA vol. 9, p. 312. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Agrimonieae > Agrimonia Rosaceae > subfam. Rosoideae
Sibling taxa
A. gryposepala, A. microcarpa, A. parviflora, A. pubescens, A. rostellata, A. striata
Subordinate taxa
Acaena, Agrimonia, Poteridium, Poterium, Sanguisorba
Name authority Torrey & A. Gray: Fl. N. Amer., 430. (1840) Lamarck & de Candolle: Syn. Pl. Fl. Gall., 333. (1806)
Web links