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annual agoseris, annual false dandelion, mountain dandelion

Photo is of parent taxon

annual agoseris

Habit Annuals.
Stems

0 or 1 (erect, 0–5 cm).

0 or 1 (internodes much shorter than subtending leaves).

Leaves

mostly erect, sometimes prostrate;

petioles not purplish, margins glabrous or ciliate;

blades usually oblanceolate to spatulate, rarely linear, 1–25 cm, margins entire or lobed;

lobes 2–3 pairs, linear to spatulate, spreading to antrorse, lobules mostly 0, glabrous or densely hairy.

erect to spreading;

blades (1–)3–15(–23) cm × (0.5–)2–10(–16) mm, mostly toothed, rarely lobed, lobes or teeth in 2–3 pairs, faces glabrous or densely, ± uniformly hairy.

Peduncles

elongating after flowering, 3–60 cm in fruit, glabrous or glabrate, or basally puberulent and apically hairy to tomentose, sometimes stipitate-glandular.

6–53 cm, lengths mostly (0.5–)1.5–4.5 times leaves at flowering, (1.5–)2–5 times leaves in fruit, glabrous or glabrate, or apically puberulent to villous.

Involucres

cylindric to hemispheric, 1–2 cm in fruit.

Receptacles

epaleate.

Florets

5–100(–300);

corollas yellow, tubes 1–5 mm, ligules 2–15 × 1–3 mm;

anthers 1–4 mm.

(5–)20–50(–100);

corollas ± equaling phyllaries at flowering, tubes 1–4 mm, ligules 2–6 × 1–2 mm;

anthers 1–1.5 mm.

Phyllaries

in 2–3 series, green or medially rosy purple, sometimes purple-black spotted or tipped, subequal to unequal, margins glabrous or ciliate, faces usually puberulent to villous, mostly stipitate-glandular, sometimes glabrous;

outer erect or spreading, adaxially usually villous to lanate, sometimes glabrous;

inner erect, ± elongating after flowering.

green or medially rosy purple, sometimes spotted, margins glabrous or ciliate, faces mostly hairy, occasionally glabrous, or villous, stipitate-glandular, translucent, yellowish or purple-septate and often purple-tipped trichomes, or eglandular with whitish opaque trichomes;

outer erect to spreading, adaxially glabrous or puberulent and eglandular.

Heads

2–10(–12) mm wide at flowering.

Cypselae

7–16 mm, bodies mostly fusiform to obconic, sometimes tumid, 2–5(–10) mm, beaks 5–11 mm, lengths 1–4 times bodies, ribs 0 or alate, straight to strongly undulate, uniform or diminishing proximally;

pappus bristles in 2–3 series, 4–9 mm.

outermost strongly differing from inner, highly variable in color, shape, ornamentation, and pubescence;

ribs flattened to alate, or 0, straight to strongly undulate, not diminishing proximally.

2n

= 18, 36.

Agoseris heterophylla

Agoseris heterophylla var. heterophylla

Phenology Flowering Mar–Sep.
Habitat Wet to dry, mostly seasonal habitats in deserts, grasslands, chaparral, steppe, oak woodlands, and open pine forests, often disturbed sites
Elevation 0–2300 m (0–7500 ft)
Distribution
from FNA
AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; nw Mexico (including Guadalupe Island) [Introduced in Europe (Sweden)]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; Mexico (Baja California, Guadalupe Island) [Introduced, Europe (Sweden)]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variety heterophylla is the most widespread of the varieties. It exhibits remarkable variation in cypsela morphology. K. L. Chambers (1963b) demonstrated that this small-flowered variety is strongly autogamous and does not outcross. The same breeding system may be present in other Agoseris taxa that exhibit reduced corolla and anther size (e.g., A. apargioides var. maritima, A. grandiflora, A. aurantiaca). Hybridization does not appear to be common; apparent intermediates with A. apargioides and A. hirsuta are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ligules 10–15 mm, much surpassing phyllaries; anthers 2–4 mm; leaf blades toothed to lobed, lobes mostly 3–4(–5) pairs
var. cryptopleura
1. Ligules 2–4 mm, subequaling phyllaries; anthers 1–1.5 mm; leaf margins entire, toothed, or lobed, lobes mostly 2–3 pairs
→ 2
2. Peduncle lengths 0.5–3 times leaves at flowering; leaf blades glabrous abaxially, pubescent adaxially; peduncles mostly glabrate, or apically tomentose
var. quentinii
2. Peduncle lengths mostly 1.5–4.5 times leaves at flowering; leaf blades uniformly glabrous or hairy; peduncles ± glabrate, or apically hairy to villous, sometimes glabrous
var. heterophylla
Source FNA vol. 19, p. 332. FNA vol. 19, p. 333.
Parent taxa Asteraceae > tribe Cichorieae > Agoseris Asteraceae > tribe Cichorieae > Agoseris > Agoseris heterophylla
Sibling taxa
A. apargioides, A. aurantiaca, A. glauca, A. grandiflora, A. hirsuta, A. monticola, A. parviflora, A. retrorsa, A. ×elata
A. heterophylla var. cryptopleura, A. heterophylla var. quentinii
Subordinate taxa
A. heterophylla var. cryptopleura, A. heterophylla var. heterophylla, A. heterophylla var. quentinii
Synonyms Macrorhynchus heterophyllus A. heterophylla var. glabra, A. heterophylla subsp. glabrata, A. heterophylla subsp. normalis, Troximon heterophyllum var. cryptopleuroides
Name authority (Nuttall) Greene: Pittonia 2: 178. (1891) unknown
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