Agoseris heterophylla |
Agoseris heterophylla var. heterophylla |
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annual agoseris, annual false dandelion, mountain dandelion |
annual agoseris |
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Habit | Annuals. | |||||||||
Stems | 0 or 1 (erect, 0–5 cm). |
0 or 1 (internodes much shorter than subtending leaves). |
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Leaves | mostly erect, sometimes prostrate; petioles not purplish, margins glabrous or ciliate; blades usually oblanceolate to spatulate, rarely linear, 1–25 cm, margins entire or lobed; lobes 2–3 pairs, linear to spatulate, spreading to antrorse, lobules mostly 0, glabrous or densely hairy. |
erect to spreading; blades (1–)3–15(–23) cm × (0.5–)2–10(–16) mm, mostly toothed, rarely lobed, lobes or teeth in 2–3 pairs, faces glabrous or densely, ± uniformly hairy. |
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Peduncles | elongating after flowering, 3–60 cm in fruit, glabrous or glabrate, or basally puberulent and apically hairy to tomentose, sometimes stipitate-glandular. |
6–53 cm, lengths mostly (0.5–)1.5–4.5 times leaves at flowering, (1.5–)2–5 times leaves in fruit, glabrous or glabrate, or apically puberulent to villous. |
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Involucres | cylindric to hemispheric, 1–2 cm in fruit. |
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Receptacles | epaleate. |
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Florets | 5–100(–300); corollas yellow, tubes 1–5 mm, ligules 2–15 × 1–3 mm; anthers 1–4 mm. |
(5–)20–50(–100); corollas ± equaling phyllaries at flowering, tubes 1–4 mm, ligules 2–6 × 1–2 mm; anthers 1–1.5 mm. |
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Phyllaries | in 2–3 series, green or medially rosy purple, sometimes purple-black spotted or tipped, subequal to unequal, margins glabrous or ciliate, faces usually puberulent to villous, mostly stipitate-glandular, sometimes glabrous; outer erect or spreading, adaxially usually villous to lanate, sometimes glabrous; inner erect, ± elongating after flowering. |
green or medially rosy purple, sometimes spotted, margins glabrous or ciliate, faces mostly hairy, occasionally glabrous, or villous, stipitate-glandular, translucent, yellowish or purple-septate and often purple-tipped trichomes, or eglandular with whitish opaque trichomes; outer erect to spreading, adaxially glabrous or puberulent and eglandular. |
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Heads | 2–10(–12) mm wide at flowering. |
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Cypselae | 7–16 mm, bodies mostly fusiform to obconic, sometimes tumid, 2–5(–10) mm, beaks 5–11 mm, lengths 1–4 times bodies, ribs 0 or alate, straight to strongly undulate, uniform or diminishing proximally; pappus bristles in 2–3 series, 4–9 mm. |
outermost strongly differing from inner, highly variable in color, shape, ornamentation, and pubescence; ribs flattened to alate, or 0, straight to strongly undulate, not diminishing proximally. |
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2n | = 18, 36. |
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Agoseris heterophylla |
Agoseris heterophylla var. heterophylla |
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Phenology | Flowering Mar–Sep. | |||||||||
Habitat | Wet to dry, mostly seasonal habitats in deserts, grasslands, chaparral, steppe, oak woodlands, and open pine forests, often disturbed sites | |||||||||
Elevation | 0–2300 m (0–7500 ft) | |||||||||
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; nw Mexico (including Guadalupe Island) [Introduced in Europe (Sweden)]
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AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; Mexico (Baja California, Guadalupe Island) [Introduced, Europe (Sweden)] |
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Discussion | Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variety heterophylla is the most widespread of the varieties. It exhibits remarkable variation in cypsela morphology. K. L. Chambers (1963b) demonstrated that this small-flowered variety is strongly autogamous and does not outcross. The same breeding system may be present in other Agoseris taxa that exhibit reduced corolla and anther size (e.g., A. apargioides var. maritima, A. grandiflora, A. aurantiaca). Hybridization does not appear to be common; apparent intermediates with A. apargioides and A. hirsuta are known. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 332. | FNA vol. 19, p. 333. | ||||||||
Parent taxa | Asteraceae > tribe Cichorieae > Agoseris | Asteraceae > tribe Cichorieae > Agoseris > Agoseris heterophylla | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Macrorhynchus heterophyllus | A. heterophylla var. glabra, A. heterophylla subsp. glabrata, A. heterophylla subsp. normalis, Troximon heterophyllum var. cryptopleuroides | ||||||||
Name authority | (Nuttall) Greene: Pittonia 2: 178. (1891) | unknown | ||||||||
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